
Belongs within: Epedanoidea.
Contains: Erecananinae, Podoctinae, Ibalonius.
The Podoctidae are an Old World tropical family of harvestmen with eyes not on a common ocularium, conspicuous spines and tubercles on the dorsum, and a long basichelicerite in males.
Podoctidae
|--+--ErecananinaeKP-GP19
| `--PodoctinaeKP-GP19
`--Ibaloniinae [Ibaloninae]SG11
|--Santobius annulipes Sørensen 1886KP-GP19, SG11
|--IbaloniusS77a
|--Serratobunus Roewer 1915R23
|--Reclinobunus Roewer 1915R23
|--Ibaloniellus Roewer 1923R23
| `--*I. karschi (Loman 1902) [=Ibalonius karschi, Paribalonius karschi]S92
|--Ceylonositalces Özdikmen 2006O06 [=Eusitalces Roewer 1915 non Brunner 1911R23]
| `--*C. parvulus (Roewer 1915) [=*Eusitalces parvulus]O06
|--Paramesoceras Roewer 1915R23
| `--*P. novoguineense Roewer 1915 [=P. novoguineensis]KM20
|--Heteropodoctis Roewer 1912R23
| `--H. quinquespinosus (Roewer 1911) [=Podoctis quinquespinosus]GG47
|--Holozoster Loman 1902R23
| `--*H. ovalis Loman 1902S92
|--Proholozoster Roewer 1915R23
| `--P. neoguinensis Roewer 1915R23
|--Ibantila Šilhavý 1969K03
| `--*I. cubana Šilhavý 1969K03
|--Paribalonius Roewer 1912R23
| `--*P. inscriptus (Loman 1902) (see below for synonymy)S92
|--Austribalonius Forster 1955F55
| `--*A. horridus Forster 1955F55
|--Heteroibalonius Goodnight & Goodnight 1947GG47
| `--*H. malkini Goodnight & Goodnight 1947GG47
|--Orobunus Goodnight & Goodnight 1947GG47
| `--*O. quadrispinosus Goodnight & Goodnight 1947GG47
|--Sitalcicus Roewer 1923R23
| |--*S. gardineri (Hirst 1911) [=Sitalces gardineri]S92
| `--S. akiyamae (Hirst 1911) [=Sitalces akiyamae]R23
|--Bonea Roewer 1913 (see below for synonymy)S77a
| |--*B. sarasinorum Roewer 1913S77a
| |--B. cippatus (Roewer 1927) [=Zmissolus cippatus]S77a
| `--B. longipalpis Suzuki 1977SG11
`--Ibalonianus Roewer 1923R23
|--I. impudens Roewer 1923R23
|--I. kueckenthali (Hirst 1912) [=Ibalonius kueckenthali]R23
`--I. quadriguttatus (Hirst 1912) [=Ibalonius quadriguttatus]R23
Podoctidae incertae sedis:
Borneojapetus Özdikmen 2006 [=Japetus Roewer 1949 non Distant 1883]O06
`--*B. longipes (Roewer 1949) [=*Japetus longipes]O06
Tandikudius rugosusGH07
Vandaravua carliGH07
Laponcea Roewer 1936S92
`--*L. cippata Roewer 1936S92
Peromona Roewer 1949S92
`--*P. erinacea Roewer 1949S92
Tryssetus Roewer 1936S92
`--*T. spinarmatus Roewer 1936S92
MelanoctisGR11
Waigeucola Roewer 1949P-G11
`--*W. palpalis Roewer 1949P-G11
Podoctops Roewer 1949KM20
`--*P. multimaculatus Roewer 1949 [=P. multimaculatum]KM20
Dimesoceras Roewer 1949S92
`--*D. lomani (Hirst 1911) [=Ibalonius lomani, Mesoceras lomani]S92
Sitalces Simon 1879S92
|--*S. novemtuberculatus Simon 1879S92 [=Sitalcicus novemtuberculatusR23]
|--S. horridusH11
|--S. mordaxH11
`--S. typusH11
Trimesoceras Roewer 1949S92
`--*T. flavopictum (Hirst 1911) [=Ibalonius flavopictus, Mesoceras flavopictum]S92
Bonea Roewer 1913 [incl. Kappacola Roewer 1949, Nurullus Roewer 1949, Parabonea Roewer 1949, Posissus Roewer 1949, Suraplus Rower 1949, Zmissolus Roewer 1927]S77
*Paribalonius inscriptus (Loman 1902) [=Ibalonius inscriptus; incl. I. bimaculatus Loman 1902, Paribalonius bimaculatus, P. bimaculatus duplex Roewer 1923]S92
*Type species of generic name indicated
References
[F55] Forster, R. R. 1955. Further Australian harvestmen (Arachnida: Opiliones). Australian Journal of Zoology 3 (3): 354–411.
[GH07] Gnaspini, P., & M. R. Hara. 2007. Defense mechanisms. In: Pinto-da-Rocha, R., G. Machado & G. Giribet (eds) Harvestmen: The Biology of Opiliones pp. 374–399. Harvard University Press: Cambridge (Massachusetts).
[GR11] Gnaspini, P., & G. C. S. Rodrigues. 2011. Comparative study of the morphology of the gland opening area among Grassatores harvestmen (Arachnida, Opiliones, Laniatores). Journal of Zoological Systematics and Evolutionary Research 49 (4): 273–284.
[GG47] Goodnight, C. J., & M. L. Goodnight. 1947. Report on a collection of phalangids from New Guinea. Transactions of the American Microscopical Society 66 (4): 328–338.
[H11] Hirst, S. 1911. On some new Opiliones from Japan and the Loo-Choo Islands. Annals and Magazine of Natural History, series 8, 8: 625–636.
[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.
[KM20] Kury, A. B., A. C. Mendes, L. Cardoso, M. S. Kury & A. A. Granado. 2020. WCO-Lite: Online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0—Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018. Published by the author: Rio de Janeiro.
[KP-GP19] Kury, A. B., A. Pérez-González & D. N. Proud. 2019. A new Indo-Malayan family of Grassatores (Arachnida: Opiliones: Laniatores). Invertebrate Systematics 33: 892–906.
[O06] Özdikmen, H. 2006. Nomenclatural changes for some Laniatores (Opiliones) genera: new substitute names and new combinations. Mun. Ent. Zool. 1 (1): 63–68.
[P-G11] Pérez-González, A. 2011. New familial assigment for two harvestman species of the infraorder Grassatores (Arachnida: Opiliones: Laniatores). Zootaxa 2757: 24–28.
[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.
[SG11] Sharma, P. P., & G. Giribet. 2011. The evolutionary and biogeographic history of the armoured harvestmen—Laniatores phylogeny based on ten molecular markers, with the description of two new families of Opiliones (Arachnida). Invertebrate Systematics 25: 106–142.
[S92] Staręga, W. 1992. An annotated check-list of Afrotropical harvestmen, excluding the Phalangiidae (Opiliones). Annals of the Natal Museum 33 (2): 271–336.
[S77] Suzuki, S. 1977. Report on a collection of opilionids from the Phillippines. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology) 27 (1): 1–120.