Belongs within: Oligoarthra.
Contains: Leptastacus, Paraleptastacus, Psammastacus, Leptocaris, Amphiascopsis, Amphiascus, Mesamphiascus, Amphiascoides, Robertsonia, Schizopera, Arenopontia, Psammopsyllinae, Stenocaris, Phyllopodopsyllus, Parastenocaris, Forficatocaris, Remaneicaris, Cletodes, Diosaccinae, Stenhelia, Bryocamptus, Attheyella, Elaphoidella, Moraria, Nitocrella, Parapseudoleptomesochra, Nitokra, Stygonitocrella, Laophonte.
The Podogennonta is a major clade of harpacticoid copepods, many of which are interstitial in habitat. Members of the clade are divided between numerous families, some of which (such as Parastenocarididae and Canthocamptidae) are primarily freshwater whereas others (such as Ameiridae and Laophontidae) are predominantly marine. The Laophontidae have a distinctive first leg structure with the two-segmented endopod longer than the one- to three-segmented exopod and ending in a protruding claw.
Synapomorphies (from Seifried 2003): Females with endopod of mandible with 3 proximal lateral setae, 3 more distal lateral setae and 3 + 2 + 2 apical setae; endopod of maxillule with 4 setae; syncoxa of maxilla with (3 + 3), 3, 3 setae, the two proximal endites fused, distal praecoxal endite with 2 flagellate setae; both coxal endites with setae of characteristic shape: anteriorly 1 claw-like seta, posteriorly 2 strong, mostly unilaterally spinulose setae with subterminal flagellum; acessory armature of fused endopod segment: 1 seta on posterior surface, seta 9 displaced to proximal segment of endopod, seta 10 lacking; endopod 3-segmented with armature formula: 2, 1, 3; setae 4 and six lost. Maxilliped 4-segmented with praecoxa and coxa separated, basis, and 1-segmented endopod; endopodal setae 1 to 4 only represented by a group of 4 small setae on anterior surface of endopod, no setae geniculated. Leg 1 of characteristic shape: endopod-1 elongate, endopod-2 and endopod-3 short; endopod-3 with 1 innter seta, terminally with anterior claw, 1 middle geniculate seta and 1 posterior miniaturised seta; 1 inner seta lacking; exopod-3 terminally with 2 geniculated setae, 2 inner setae lacking. Formula of armature of leg 1: coxa 0-0, basis I-I, exopod I-0, I-1, III-2-0, endopod 0-1, 0-1, I-2-1. Leg 5 with endopodal lobe with six setae: 3 inner setae, 2 terminal setae, 1 outer seta; exopod foliated with 8 setae: 3 inner, 2 terminal, 3 outer setae.
Podogennonta
|--ArgestidaeS03
|--AdenopleurellidaeS03
|--CancrincolidaeS03
|--Cletopsyllus Willey 1935N41b [CletopsyllidaeS03, Cletopsyllinae]
|--CristacoxidaeS03
|--Normanella Brady 1880N41b [NormanellidaeS03, Normanellinae]
|--‘Stenhelia’ glacialis Brady 1918N41b [PseudotachidiidaeS03, Pseudotachidiinae]
|--Ismardis Leigh-Sharpe 1936 [Ismardiidae]S03
|--ThompsonulaN41b [ThompsonulidaeS03]
| `--T. hyaenae (Thompson 1889) [incl. Robertsonia aculeifera Klie 1913]N41b
|--BalaenophilusW81 [BalaenophilidaeS03]
| `--B. unisetusW81
|--Lourinia Wilson 1924N41a (see below for synonymy)
| `--L. armata (Claus 1866)W67 (see below for synonymy)
|--Laophontopsis Sars 1908N41b [=Cleta Claus 1863 (preoc.)N41b; LaophontopsidaeS03]
| |--*L. lamellifera (Claus 1863) [=*Cleta lamellifera]N41b
| `--L. secunda (Sewell 1924)N41b
|--Parastenhelia Thompson & Scott 1903N41b [incl. MicrothalestrisN41b; ParastenheliidaeS03]
| |--‘Microthalestris’ forficulaB26
| |--P. hornelli Thompson & Scott 1903W67
| `--P. spinosa (Fischer 1860)W67
|--Rhynchothalestris [Rhynchothalestridae]S03
| |--R. helgolandica (Claus 1863)S03
| |--R. rufocinctaN44
| `--R. vanhoeffeni Brady 1910N44
|--Laophontodes Scott 1894N41b [AncorabolidaeS03]
| |--L. armatus (Lang 1936)W67 [incl. Laophonte echinata Willey 1930N41b]
| |--L. bicornis Scott 1896N44
| |--L. brevis Nicholls 1944N44
| `--L. typicusN44
|--HuntemanniidaeS03
| |--Huntemannia Poppe 1884S03
| `--NannopusS03
| |--N. abyssi Sars 1921S03
| `--N. palustris Brady 1880S03
|--DactylopusiaN41b [DactylopusiidaeS03]
| |--D. micronyxN41b
| |--D. oculata Gurney 1927N41b
| |--D. platysoma Thompson & Scott 1903N41a
| |--D. tisboidesN41b
| `--D. vulgarisN44
|--Delamarella Chappuis 1954W86 [LatiremidaeS03]
| |--D. arenicola Chappuis 1954W86
| |--D. eximia (Božić 1969)W86
| |--D. galateae Cottarelli 1971W86
| |--D. karamani Petkovski 1957W86
| `--D. phyllosetosa Kunz 1984W86
|--PhyllognathopodidaeS03
| |--Allophyllognathopus Kiefer 1967R86
| | `--A. brasiliensis Kiefer 1967R86
| `--Phyllognathopus Mrazek 1893R86
| |--P. bassoti Rouch 1972R86
| `--P. viguieri (Maupas 192)B01
|--Leptastacidae [Leptastacinae]S03
| |--LeptastacusW86
| |--ParaleptastacusW86
| |--PsammastacusW86
| `--Arenocaris Nicholls 1935W86
| `--A. bifida Nicholls 1935W86
|--Orthopsyllus Brady & Robertson 1873N41a [OrthopsyllidaeS03]
| |--O. linearis (Claus 1866)W67
| |--O. littoralis Nicholls 1942N42b
| |--O. major Klie 1939N41a
| |--O. propinquus Monard 1926N42b
| |--O. rugosus Nicholls 1941N42b
| |--O. similis Nicholls 1942N42b
| `--O. wallini Lang 1934N42b
|--MetidaeS03
| |--LaubieriaW67
| | |--*L. corallicola Soyer 1966W67
| | `--L. secunda Wells 1967W67
| `--Metis Philippi 1843N41a
| |--M. holothuriae (Edwards 1891)W67 [=Abacola holothuriaeB26]
| |--M. ignea (see below for synonymy)B26
| `--M. jousseaumei (Richard 1892) [=Ilyopsyllus jousseaumei]N42b
|--DarcythompsoniidaeS03
| |--Kristensenia Por 1983S03
| |--LeptocarisS03
| |--DarcythompsoniaW86
| `--HorsiellaN44
| |--H. brevicornisN44
| |--H. gurneyi Nicholls 1944N44
| `--H. trisetosa Kunz 1935N44
|--MiraciidaeKM13
| | i. s.: Eoschizopera Wells & Rao 1976KM13
| `--AmphiascinaeN41b
| |--AmphiascopsisN41b
| |--AmphiascusN41b
| |--MesamphiascusN41b
| |--AmphiascoidesN41b
| |--RobertsoniaN41b
| `--SchizoperaKM13
|--Leptopontiidae [Leptopontiinae]S03
| | i. s.: ArenopontiaW86
| | Notopontia Bodiou 1977W86
| | |--N. galapagoensis Mielke 1982W86
| | `--N. stephanieae Bodiou 1977W86
| | Ichnusella Cottarelli 1971GL04
| | |--I. eione Cottarelli 1971R86
| | `--I. pasquinii (Cottarelli 1969)R86
| | Parasewellina Cottarelli, Saporito & Puccetti 1986GL04
| |--PsammopsyllinaeGL04
| `--Leptopontia Scott 1902 [Leptopontiinae]S03
| |--L. curvicauda Scott 1902W86
| `--L. punctata Huys & Conroy-Dalton 1996S03
|--Cylindropsyllidae [Cylindropsyllinae]S03
| |--StenocarisW86
| |--Boreopontia Willems 1981W86
| | `--B. heipi Willems 1981W86
| |--Cylindropsyllus Brady 1880W86
| | |--C. laevis Brady 1880W86
| | `--C. remanei Kunz 1949W86
| |--Syrticola Willems & Claeys 1982W86
| | |--S. flandricus Willems & Claeys 1982W86
| | `--S. galapagoensis (Mielke 1982)W86
| `--Evansula Scott 1906W86
| |--E. arenicola Nicholls 1939W86
| |--E. incerta (Scott 1892)W86
| `--E. pygmaea (Scott 1903)W86
|--Rhizothrix Brady & Robertson 1875W86 [RhizothricidaeS03]
| | i. s.: R. curvata Brady & Robertson 1880W86
| | R. gracilis (Scott 1903)W86
| | R. minuta (Scott 1903)W86
| | R. reducta Noodt 1952W86
| | |--R. r. reductaW86
| | `--R. r. noodti Galhano 1970W86
| | R. scotti Lang 1936W86
| | R. spinosa Coull 1971W86
| | R. tenella (Wilson 1932)W86
| | R. wilsoni Bodin 1977W86
| `--R. (Rhizothrix)W67
| |--R. (R.) pubescens Por 1959W67
| `--R. (R.) quadriseta Wells 1967W67
|--Tetragonicipitidae [Pteropsyllidae, Tetragonicepsidae]S03
| |--PhyllopodopsyllusPH13
| |--DiagonicepsN44
| |--Laophontella Thompson & Scott 1903W67, N41b
| | `--L. armata (Willey 1935) [=Phyllopodopsyllus armatus]N44
| |--TetragonicepsW67
| | |--T. bergensisW67
| | |--T. brownei Wells 1967W67
| | `--T. scottiW67
| `--PteropsyllusW67
| |--P. consimilis (Scott 1894)W67
| `--P. plebius Monard 1935W67
| |--P. p. plebiusW67
| `--P. p. furcatus Kunz 1938W67
|--Parastenocarididae [Fontinalicaridinae, Parastenocaridinae]S03
| |--ParastenocarisCMA05
| |--ForficatocarisCMA05
| |--RemaneicarisCMA05
| |--Cafferocaris Jakobi 1972GL04
| |--Paraforficatocaris Jakobi 1972R86
| | `--P. paranaensis Jakobi 1972R86
| |--Murunducaris Reid 1994CMA05
| | `--M. juneae Reid 1994CMA05
| |--Potamocaris Dussart 1970CMA05
| | |--P. bidens (Noodt 1955)GL04
| | |--P. bifida Dussart 1979R86
| | `--P. estevesi Reid 1991GL04
| `--Simplicaris Galassi & De Laurentiis 2004CMA05
| |--*S. lethaea Galassi & De Laurentiis 2004FS14
| |--‘Parastenocaris’ aedes Hertzog 1938 [=Pannonicaris aedes]GL04
| |--‘Parastenocaris’ hippuris Hertzog 1938GL04
| `--S. veneris (Cottarelli & Maiolini 1980) [=Parastenocaris veneris]GL04
|--Cletodidae [Cletodoidea]S03
| |--CletodesN41b
| |--Parepactophanes Kunz 1935W86
| | |--P. dubia Noodt 1958W86
| | `--P. minuta Kunz 1935W86
| |--Cletocamptus Schmankevitch 1875R86
| | |--C. albuquerquensisG88
| | |--C. deitersi (Richard 1897)S-MR03
| | `--C. xenuus Por 1968R86
| |--Tryphoema Monard 1926W86
| | |--T. bocqueti (Božić 1954)W86
| | |--T. lusitanica (Wells & Clark 1965)W86
| | |--T. porca Monard 1926W86
| | |--T. ramabula (Pennak 1942)W86
| | |--T. riedli Coull 1971W86
| | `--T. scilloniensis (Wells 1968)W86
| |--ParanannopusW56
| `--EnhydrosomaW86
| |--E. barnishi Wells 1967W67
| |--E. buchholtzi (Boeck 1872)W67
| |--E. curticaudatumN41b
| |--E. curvirostreW67
| |--E. latipes (Scott 1909)W67
| |--E. littorale Wells 1967W67
| |--E. longifurcatum Sars 1909W67
| `--E. parapropinquum Gomez 2003S03
|--HarpacticidaeS03
| |--HarpacticellaSS03
| | |--H. inopinataB50
| | |--H. lacustrisB50
| | `--H. paradoxaB50
| |--Perissocope litoralis Lang 1934S03
| |--Zausopsis mirabilis Lang 1934S03
| |--ZausodesS03
| | |--Z. septimus Lang 1965S03
| | `--Z. sextus Lang 1965S03
| |--TigriopusSS03
| | |--T. brevicornisCCG01
| | |--T. californicusSC17
| | |--T. fulvusW67 [=Harpacticus fulvusPP64]
| | `--T. japonicusWS10
| |--HarpacticusS03
| | |--H. chelifer (Müller 1776)W67
| | |--H. compsonyx Monard 1926W67
| | |--H. gracilis Claus 1863W67
| | |--H. littoralisB26
| | |--H. trisetosus Lang 1948S03
| | `--H. uniremisB26
| `--Zaus Goodsir 1845S03, N42a
| |--Z. abbreviatus Sars 1911N42a
| |--Z. aurelii Poppe 1884 [incl. Z. caeruleus Campbell 1929, Z. intermedius Nicholls 1939]N42a
| |--Z. contractus Thomson 1883N42a
| |--Z. goodsiri Brady 1880N42a
| |--Z. sarsi Nicholls 1942N42a
| `--Z. spinatus Goodsir 1845N42a
|--ThalestridaeS03
| | i. s.: Amenophia Boeck 1865S03
| | Paradactylopodia brevicornis (Claus 1866)W67
| | DiarthrodesW67
| | |--D. dissimilisW67
| | |--D. glaber Wells 1967W67
| | |--D. minutus (Claus 1863)W67
| | |--D. nanus (Scott 1914)W67
| | `--D. sarsi (Scott 1909)W67
| | Idomene [incl. Xouthous]W67
| | |--I. aberrans Por 1964W67
| | |--I. aemula Thompson & Scott 1903W67
| | |--I. laticaudata (Thompson & Scott 1903)W67
| | |--‘Xouthous’ maldivae Sewell 1940W67
| | `--I. simulans (Brady 1910)W67
| | Dactylopodella flavaN41b
| |--DactylopodiinaeN41a
| | |--Dactylopodia tisboides (Claus 1863)W67
| | `--Eudactylopus Scott 1909N41a
| | |--E. australis Nicholls 1941N42b
| | |--E. latipes (Scott 1894)N41a
| | |--E. robustus (Claus 1863)W67
| | `--E. spectabilis (Brian 1923)N41a
| `--ThalestrinaeN41a
| |--Thalestris longimanaBK77
| `--Phyllothalestris Sars 1905N41a
| |--P. lata Nicholls 1942N42b
| |--P. mysis (Claus 1863)N41a
| | |--P. m. mysisN42b
| | `--P. m. harringtoni Willey 1935N42b
| |--P. paramysis Monard 1928N42b
| |--P. royi (Monard 1928)N42b
| `--P. sarsi Sewell 1940W67
|--DiosaccidaeS03
| | i. s.: Pseudamphiascopsis ismaelensis (Monard 1936)W67
| | Bulbamphiascus imus (Brady 1872)W67
| | Haloschizopera junodi (Monard 1935)W67
| | Paramphiascella robinsoni (Scott 1902)W67
| | MelimaW67
| | HelmutkunziaW86
| | Typhlamphiascus Lang 1948R86
| | `--T. latifurcata Por 1968R86
| | Pseudostenhelia Wells 1967W67
| | `--*P. prima Wells 1967W67
| | Paramphiascoides Wells 1967W67
| | `--*P. mixtus Wells 1967W67
| | Actopsyllus Wells 1967W67
| | `--*A. longipes Wells 1967W67
| | Schizoperoides Por 1968R86
| | `--S. expeditionis Por 1968R86
| | Goffinella Wilson 1932W86
| | `--G. stylifer Wilson 1932W86
| | Balucopsylla Rao 1972W86
| | `--B. similis Rao 1972W86
| | Protopsammotopa Geddes 1968W86
| | |--P. norvegica Geddes 1968W86
| | `--P. wilsoni Wells 1977W86
| | RobertgurneyaW67
| | |--R. rostrata (Gurney 1927)W67
| | |--R. similis (Scott 1896)W67
| | `--R. spinulosa (Sars 1911)W67
| | Psammotopa Pennak 1942W86
| | |--P. chappuisi Noodt 1955W86
| | |--P. phyllosetosa (Noodt 1952)W86
| | |--P. polyphylla Noodt 1955W86
| | `--P. vulgaris Pennak 1942W86
| |--DiosaccinaeN41b
| `--StenheliinaeN41b
| |--StenheliaN41b
| `--Pseudomesochra Scott 1902 [incl. Stenheliopsis Sars 1906]N41b
|--CanthocamptidaeS03
| |--Nannomesochra arupinensis (Brian 1925)W67
| |--BryocamptusS-MR03
| |--AttheyellaS-MR03
| |--ElaphoidellaKM13
| |--MorariaS-MR03
| |--Australocamptus hamondi Karanovic 2004KM13
| |--Gulcamptus Miura 1969R86
| | `--G. uenoi Miura 1969R86
| |--Antarctobiotus Chappuis 1930R86
| | `--A. bahamondei Ebert & Noodt 1975R86
| |--Epactophanoides Borutzky 1966R86
| | `--E. udegeicus Borutzky 1966R86
| |--Stygepactophanes Moeschler & Rouch 1984R86
| | `--*S. jurassicus Moeschler & Rouch 1984R86
| |--Echinocamptus Chappuis 1928R86
| | |--E. echinatus (Mrázek 1893) [=Bryocamptus (Echinocamptus) echinatus]FS14
| | `--E. georgevitchi (Chappuis 1924)B01
| |--‘Paracamptus’ Chappuis 1929 non Casey 1895R86
| | |--P. baikalensisW56
| | |--P. gasparoi Stoch 1998B01
| | |--P. nakamurai Chappuis 1958R86
| | |--P. reductus Wilson 1956W56
| | `--P. schmeiliW56
| |--Morariopsis Borutzky 1931R86
| | |--M. kieferi Petkovski 1959R86
| | `--M. scotenophila (Kiefer 1930)B01
| |--Canthocamptus Westwood 1836R86
| | |--C. insoletus Chappuis 1928R86
| | |--C. morimotoi Miura 1969R86
| | `--C. staphylinusD01
| |--Spelaeocamptus Chappuis 1933R86
| | |--S. incertus Petkovski 1956R86
| | |--S. neotropicus Noodt 1963R86
| | `--S. spelaeus (Chappuis 1925)R86
| |--Mesochra Boeck 1865R86, S03
| | |--M. hiunumaensis Kikuchi 1972R86
| | |--M. lilljeborgi [incl. Laophonte hecate Brehm 1910]N41b
| | |--M. pygmaea (Claus 1863)W67
| | `--M. rostrata (Gurney 1927)W67
| |--Antrocamptus Chappuis 1956R86
| | |--A. catherinae Chappuis & Rouch 1960R86
| | |--A. chappuisi Rouch 1970R86
| | |--A. coiffaiti Chappius 1956R86
| | |--A. longifurcatus Rouch 1970R86
| | `--A. stygius Rouch 1970R86
| |--Maraenobiotus Mrazek 1893R86
| | |--M. bruceiR86
| | | |--M. b. bruceiR86
| | | `--M. b. carpathicus Chappuis 1928R86
| | `--M. insignipesR86
| | |--M. i. insignipesR86
| | `--M. i. kyzylkumicus Borutzky 1972R86
| `--Ceuthonectes Chappuis 1924R86
| |--C. chappuisi Rouch 1980R86
| |--C. gallicus Chappuis 1928R86
| |--C. hungaricus Ponyi 1958R86
| |--C. mirabilis Miura 1964R86
| |--C. rouchi Petkovski 1984R86
| |--C. serbicus Chappuis 1924 [incl. Morariodes colchidana Borutzky 1930, C. colchidanus]R86
| `--C. vievilleae Rouch 1980R86
|--AmeiridaeS03
| | i. s.: ‘Amphiascus’ mucronatus Brady 1910N41b
| | ‘Amphiascus’ proximus Scott 1914N41b
| |--Psammoleptomesochra australis (Mielke 1994) [=Paramesochra australis]S03
| |--PseudameiraW67
| |--ProameiraW67
| |--NitocrellaR86
| |--LeptomesochraW67
| |--ParapseudoleptomesochraS-MR03
| |--NitokraPH13
| |--StygonitocrellaR86
| |--SicamaeiraW86
| |--Inermipes humphreysi Lee & Huys 2002HA13
| |--Biameiropsis barrowensis Karanovic 2006HA13
| |--Pseudoleptomesochra Lang 1965W86
| | `--P. typica Lang 1965W86
| |--Psyllocamptus Scott 1899R86
| | |--P. minutus Sars 1911W67
| | `--P. monachus Chappuis 1938R86
| |--Paraleptomesochra Wells 1967W86
| | |--*P. minima Wells 1967W67
| | `--P. wellsi Rao 1972W86
| |--Philoleptomesochra Wells 1967W67
| | |--*P. reductiforma Wells 1967W67
| | `--P. secunda Wells 1967W67
| |--Karllangia Noodt 1964W86
| | |--*K. arenicola Noodt 1964W67, W86
| | |--K. psammophila Wells 1967W86
| | `--K. tertia Kunz 1975W86
| |--Sarsameira Wilson 1924 [incl. Pseudosarsameira]W67
| | |--S. exilis (Scott & Scott 1894)W67
| | |--S. minor Wells 1967W67
| | `--S. pendula (Shen & Bai 1956)W67
| |--Psammameira Noodt 1952W67
| | |--*P. hyalina Noodt 1952W67
| | |--P. grandis (Nicholls 1939) [=Ameira grandis]W67
| | `--P. reducta Wells 1967W67
| |--Nitocrellopsis Petkovski 1976R86
| | |--N. elegans (Chappuis & Rouch 1959)R86
| | |--N. intermedia (Chappuis 1937)R86
| | `--N. ioneli (Dumont & Decraemer 1974)R86
| |--Praeleptomesochra Lang 1965W86
| | |--P. africana (Kunz 1951)W86
| | |--P. phreatica Pesce 1981R86
| | |--P. pygmaea (Vervoort 1964)W86
| | `--P. similis Lang 1965W86
| |--AmeiraW86
| | |--A. exiguaW67
| | |--A. leptoderma (Klie 1950)W67
| | |--A. longipes Boeck 1864W67
| | |--A. parvula (Claus 1866) [=Amphiascus parvulus]N41b
| | `--A. simulansW67
| |--Pseudoleptomesochrella Lang 1965W86
| | |--P. bisetosa Lindgren 1975W86
| | |--P. halophila (Noodt 1952)W86
| | |--P. incerta (Chappuis & Delamare Deboutteville 1956)W86
| | |--P. marina (Chappuis & Rouch 1960)W86
| | `--P. pontica Apostolov 1969W86
| |--Parevansula Guille & Soyer 1966W86
| | |--P. elegans (Marinov 1974)W86
| | |--P. mediterranea Guille & Soyer 1966W86
| | |--P. reductiforma (Wells 1967)W86
| | |--P. secunda (Wells 1967)W86
| | |--P. vermiformis Moore 1976W86
| | `--P. wellsi (Marinov 1973)W86
| `--Interleptomesochra Lang 1965W86
| |--I. attenuata (Scott 1896)W86
| |--I. boguensis Lindgren 1975W86
| |--I. elongata (Božić 1955)W86
| |--I. eulitoralis (Noodt 1952)W86
| |--I. noodti Galhano 1968W86
| |--I. reducta Lang 1965W86
| `--I. tenuicornis (Sars 1911)W86
`--Laophontidae [Namakosiramiidae]S03
|--Heterolaophonte quinquespinosa (Sewell 1924)W67
|--KlieonychocamptusW67
|--Pseudocletopsyllus Vervoort 1964W67
|--Mictyricola Nicholls 1957W67
|--Pholenota Vervoort 1964W67
|--Novolaophonte viatorum Cottarelli, Saporito & Puccetti 1983W86
|--LaophonteN41b
|--Platychelipus Brady 1880N41b
|--Sarsocletodes Wilson 1924 [=Pseudocletodes Sars 1921 (preoc.), Pseudoplatychelipus Lang 1936]N41b
|--Donsiella Stephenson 1935N41b
|--Psammolaophonte Wells 1967W86
| `--*P. spinicauda Wells 1967W67
|--Apolethon Wells 1967W67
| `--*A. fumator Wells 1967W67
|--Esola Edwards 1891N41b
| `--E. spelea (Chappuis 1938)R86
|--Cubanocleta Petkovski 1977R86
| `--C. noodti Petkovski 1977R86
|--Mexicolaophonte Cottarelli 1977W86
| `--M. arganoi Cottarelli 1977W86
|--Arenolaophonte Lang 1965W86
| `--A. stygia Lang 1965W86
|--Stygolaophonte Lang 1965W86
| `--S. arenophila Lang 1965W86
|--Pseudolaophonte Scott 1896N41b
| `--P. spinosa (Thompson 1893) [incl. P. aculeata Scott 1896]N41b
|--Harrietella Scott 1906N41b
| `--H. simulans (Scott 1894)N41b
|--Hemilaophonte Jakubisiak 1932N41b
| `--H. janinae Jakubisiak 1932N41b
|--Lobitella Monard 1934N41b
| `--L. apoda Monard 1934N41b
|--OnychocamptusPH13
| |--O. bengalensis (Sewell 1934)PH13
| `--O. mohammed (Blanchard & Richard 1891)S-MR03 [=Laophonte mohammedS-MR03; incl. L. humilis Brian 1929N41b]
|--Tapholeon Wells 1967W67
| |--*T. ornatus Wells 1967W67
| `--T. uniarticulatus Wells 1967W67
|--ParalaophonteW67
| |--P. brevirostris (Claus 1863)W67
| |--P. congenera (Sars 1908)W67
| |--P. quinquespinosaP71
| `--P. tenera (Sars 1920)W67
|--Asellopsis Brady & Robertson 1873N41b
| |--A. duboscqui Monard 1926N41b
| |--A. hispida Brady & Robertson 1873N41b
| |--A. intermedia (Scott 1895)N41b
| `--A. littoralis Nicholls 1939N41b
|--Klieonychocamptoides Noodt 1958W86
| |--K. arenicola (Chappuis & Delamare Deboutteville 1956)W86
| |--K. arganoi Cottarelli & Mura 1980W86
| |--K. itoi Mielke 1981W86
| `--K. remanei Noodt 1958W86
|--Echinolaophonte Nicholls 1941N41b
| |--*E. horrida (Norman 1876) [=Laophonte horrida]N41b
| |--E. armiger (Gurney 1927) [=Laophonte armiger; incl. L. hystrix Brian 1928, L. steueri van Douwe 1929]N41b
| |--E. brevispinosa (Sars 1908) [=Laophonte brevispinosa]N41b
| `--E. mirabilis (Gurney 1927) [=Laophonte mirabilis]N41b
|--Laophontina Norman & Scott 1905N41b
| |--L. acantha Noodt 1955W86
| |--L. distincta Wells 1967W86
| |--L. dubia Norman & Scott 1905N41b
| |--L. noodti Kunz 1983W86
| |--L. reducta Coull & Zo 1980W86
| |--L. triarticulata Coull & Zo 1980W86
| `--L. variabilis Coull & Zo 1980W86
`--Aphrolaophonte Chappuis 1960W86
|--A. aequatorialis Cottarelli & Mura 1982W86
|--A. brevipes (Chappuis 1954)W86
|--A. leonis Cottarelli & Mura 1982W86
|--A. michaelae Cottarelli & Mura 1982W86
|--A. monodi Chappuis 1960W86
|--A. pori Masry 1970W86
|--A. renaudi (Chappuis & Delamare Deboutteville 1956)W86
`--A. schmidti Mielke 1981W86
Podogennonta incertae sedis:
Antiboreodiosaccus Lang 1944S03
Dactylopina Brady 1910S03
Flavia Brady 1899S03
Mawsonella Brady 1918S03
Pyrocletodes Coull 1973S03
Tisemus Monard 1928S03
Metahuntemannia Smirnov 1946S03
|--M. spinipes Dahms & Pottek 1992S03
`--M. triarticulata Schriever 1984S03
Schizacron vervoorti (Fiers 1987)S03
Psammonitocrella Rouch 1992GL04
Lourinia Wilson 1924N41a [=Jurinia Claus 1866 (preoc.)N41a; incl. Ceylonia Thompson & Scott 1903N41a, Ceyloniella Wilson 1924N41a; Ceyloniellidae, Louriniidae S03]
Lourinia armata (Claus 1866)W67 [=Jurina armataN41a, Ceylonia armataN41a, Ceyloniella armataN41a; incl. Ceylonia aculeata Thompson & Scott 1903N41a, Ceyloniella aculeataN41a, Ceylonia aculeata var. adriatica Brian 1923N41a, Ceyloniella aculeata var. adriaticaN41a]
Metis ignea [incl. Ilyopsyllus coriaceus Brady & Robertston 1873, I. natans Williams 1907, I. sarsi Sharpe 1911]B26
*Type species of generic name indicated
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Dear Chris, that is an Anoplosomella. It is not considered Ameiridae anymore, being transfered to Argestidae.
Thank you, Paulo, I've edited the picture caption accordingly.