Polycestinae

 Astraeus fraterculus, copyright Western Australian Museum.

Belongs within: Buprestidae.
Contains: Acmaeodera, Polycesta.

The Polycestinae are a group of jewel beetles that often have strongly punctured and/or carinate elytra and a sternal cavity for reception of the prosternal process often formed entirely from the mesosternum. The sternal cavity is partially formed by the metasternum in the Australasian genus Astraeus, members of which are generally strongly convex.

Members of the North American genus Thrincopyge have the metacoxal plates dilated medially, and a deep groove around the apical half of the last visible sternite (Bellamy & Nelson 2002).

<==Polycestinae [Acmaeoderinae, Acmaeoderites, Polycestides, Ptosimites]
    |--Pseudacherusia Kerremans 1904B85
    |--Perucola Thery 1925 [Perucolini]B85
    |--Prospheres Thomson 1878 [=Phospheres (l. c.); Prospherini]B85
    |--Nothomorpha Thomson 1878 [Nothomorphini]B85
    |--Xyroscelis Thomson 1878 [Xyroscelini]B85
    |    `--X. crocata [incl. X. melanosticta Hope 1846, X. nodosa Hope 1846]M86
    |--Thrincopyge LeConte 1858B14, B85 [Thrincopygina, Thrincopyginae, Thrincopygini]
    |    |--T. alacris LeConte 1858B14
    |    `--T. ambiensBN02
    |--TylaucheniniB85
    |    |--Tylauchenia Burmeister 1872 [incl. Ocypetes Thomson 1878 non Lesson 1842]B85
    |    |--Neocypetes Cobos 1973B85
    |    `--Ancylotela Waterhouse 1882B85
    |--AcherusiniB85
    |    |--Hayekina Cobos 1980B85
    |    |--Bulis Laporte & Gory 1837B85
    |    `--Acherusia Laporte & Gory 1837B85
    |         |  i. s.: A. childreniM86
    |         `--A. (Nelsonila Cobos 1980)B85
    |--Tyndarini [Tyndarina]B85
    |    |--Paraancylotela Cobos 1959B85
    |    |--Bordonia Cobos 1980B85
    |    |--Tyndaris Thomson 1857B85
    |    `--Paratyndaris Fisher 1919 [incl. Barberia Cobos 1980 (nom. inv.), Tucsonia Cobos 1980 (nom. inv.)]BN02
    |         |  i. s.: P. olneyaeBN02
    |         `--P. (Knulliella Cobos 1980)BN02
    |--Polyctesini [Polyctesina]B85
    |    |--Paraxenopsis Cobos 1980B85
    |    |--Xenopsis Saunders 1867B85
    |    |--Polyctesis Marseul 1865 [incl. Schoutedenia Obenberger 1924 (preoc.), Schoutendeiastes Burgeon 1941]B85
    |    |--Chrysophana LeConte 1859B85, BN02
    |    |    `--C. placidaBN02
    |    `--Beerellus Nelson 1982B85, BN02
    |         `--B. taxodiiBN02
    |--Astraeus Laporte & Gory 1837 [Astraeusini]B85
    |    |--A. aberransB14
    |    |--A. adamsiB14
    |    |--A. caledonicusB14
    |    |--A. carnabyiB14
    |    |--A. dedariensisB14
    |    |--A. flavopictusM86
    |    |--A. fraterculus van de Poll 1889B14
    |    |--A. major Blackburn 1890B90
    |    |--A. meyricki Blackburn 1890B90
    |    |--A. navarchis Thoms. 1856M86 [=Conognatha navarchisB90]
    |    |--A. samoellei Saund. 1868M86 [incl. A. mastersi Macleay 1872B90, M86, A. pygmaeusB90]
    |    `--A. tepperi Blackburn 1890B90
    |--Mastogeniini [Mastogeniina, Mastogeninae]BN02
    |    |--Helferella Cobos 1957B85
    |    |--Obenbergerietta Strand 1942 [incl. Sicardia Thery 1909 (preoc.)]B85
    |    |--Ankareus Kerremans 1894 [incl. Pseudianthe Fairmaire 1901]B85
    |    |--Siamastogenius Toyama 1983B85
    |    |--Neomastogenius Toyama 1983B85
    |    |--Exaesthetus Waterhouse 1887B85
    |    |--Trigonogya Schaeffer 1919B85
    |    |    `--T. reticulaticollis (Schaeffer 1904)BN02
    |    |--Micrasta Kerremans 1893B85
    |    |    `--M. oakleyi Fisher 1935BN02
    |    `--Mastogenius Solier 1850 [incl. Haplostethus LeConte 1859]BN02
    |         |--M. primaevusP92
    |         `--M. robustusBN02
    |--AcmaeoderiniB85
    |    |  i. s.: Microacmaeodera Cobos 1966B85
    |    |         Xantheremia Volkovitsh 1979B85
    |    |         Cochinchinula Volkovitsh 1984B85
    |    |           `--*C. quadriareolata [=Acmaeodera quadriareolata]B85
    |    |         Acmaeoderella Cobos 1955B85
    |    |           |--A. (Carininota Volkovitsh 1979)B85
    |    |           |--A. (Euacmaeoderella Volkovitsh 1979)B85
    |    |           |--A. (Kocheridia Cobos 1958)B85
    |    |           |--A. (Liogastria Volkovitsh 1979)B85
    |    |           `--A. (Omphalothorax Volkovitsh 1979)B85
    |    |--Acmaeoderoides Van Dyke 1942 [Acmaeoderoidina, Acmaeoderoidini]BN02
    |    |    `--A. knulliBN02
    |    |--AcmaeoderinaBN02
    |    |    |--AcmaeoderaB14
    |    |    |--Squamodera Nelson 1996BN02
    |    |    |--Acmaeoderopsis Barr 1974BN02
    |    |    `--Anambodera Barr 1974BN02
    |    `--Ptosimina [Ptosimini]BN02
    |         |--Paratrachys Saunders 1873 [Paratrachysae]B85
    |         `--PtosimaeB85
    |              |--Ptosima Serville in Dejean 1833BN02
    |              |    `--P. gibbicollisBN02
    |              `--Sponsor Laporte & Gory 1839B85
    |                   |--S. (Neoptosima Thery 1905)B85
    |                   `--S. (Stenianthe Fairmaire 1886)B85
    `--Polycestini [Polycestina]B85
         |--PolycestaB85
         |--Pseudocastalia Kraatz 1896B85
         |--Agenjosiana Cobos 1981B85
         |--Madecacesta Descarpentries 1975B85
         |--Cobosesta Holm 1982B85
         |--Jelinekia Cobos 1981B85
         |--Kogania Cobos 1981B85
         |--Madecastalia Descarpentries 1975B85
         |--Parapolycesta Cobos 1981B85
         |--Strigopteroides Cobos 1981B85
         |--Polycestoides Kerremans 1902B85
         |--Polycestaxis Obenberger 1920B85
         |--Polycestaxia Cobos 1981B85
         |--Pseudopolycesta Cobos 1981B85
         |--Castaliella Cobos 1981B85
         |--Micropolycesta Cobos 1981B85
         |--Aldabrica Cobos 1981B85
         |--Polycestella Kerremans 1902B85
         |--Thurntaxisia Schatzmayr 1929 [incl. Acassidiocesta Mandl 1972]B85
         |--Paracastalia Kerremans 1902 [incl. Lypocesta Cobos 1981]B85
         |    `--P. (Mattheesta Holm 1982)B85
         |--Polycestina Cobos 1981B85
         |    `--P. (Damarana Cobos 1981)B85
         |--Neopolycesta Kerremans 1904 [incl. Holmia Cobos 1981]B85
         |    `--N. (Peringueyesta Holm 1982)B85
         `--Strigoptera Dejean 1838 [incl. Castalia Gory & Laporte 1837 non Savigny 1820]B85
              |--S. bimaculata (Linnaeus 1758)B35, L58 [=Buprestis bimaculataB35, Castalia bimaculataM86; incl. C. impustulataM86]
              |--S. bipustulataB35 [=Buprestis bipustulataB35, Castalia bipustulataM86]
              |--‘Castalia’ globithorax Thoms. 1878M86
              `--‘Castalia’ pacificaM86

*Type species of generic name indicated

References

[B85] Bellamy, C. L. 1985. A catalogue of the higher taxa of the family Buprestidae (Coleoptera). Navorsinge van die Nasionale Museum Bloemfontein 4 (15): 405–472.

[BN02] Bellamy, C. L., & G. H. Nelson. 2002. Buprestidae Leach 1815. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 98–112. CRC Press.

[B90] Blackburn, T. 1890. Notes on Australian Coleoptera, with descriptions of new species. Part V. Proceedings of the Linnean Society of New South Wales, series 2, 4 (4): 1247–1276.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d’Urville. Faune entomologique de l’océan Pacifique, avec l’illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

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