Belongs within: Lycaenidae.
Contains: Theclinesthes, Nacaduba, Prosotas, Everes, Tongeia, Polyommatus, Albulina, Agriades, Lycaeides, Agrodiaetus, Cyaniris, Plebejus, Celastrina, Maculinea, Glaucopsyche.
The Polyommatini are a cosmopolitan group of butterflies. Members typically lack a basal dark streak on the underside of the fore wing except in a few species possessing tailed wings.
The tribe is divided between numerous groups with veins Sc and R1 completely free from each other in the Lampides, Leptotes, Lycaenopsis, Euchrysops, Polyommatus and Glaucopsyche groups but not in other groups. The genus Lampides is distinguished by unique, flask-shaped androconia. Leptotes has a fuscous streak on the underside of the fore wing below vein Sc. Males of the Lycaenopsis group have genitalia typically lacking brachia but with a developed coecum (Wang & Fan 2002).
Characters (Eliot 1973): Fore wing with veins 11 and 12 variable, may be free, touch or anastomose; in a few species vein 11 may arise from vein 10; an additional vein linking veins 11 and 12 rarely occurs. Hind wing tailed or tailless. Secondary sexual characters comprise various types of specialized scales, of which battledore androconia, arranged in alternate rows with ordinary scales, are much the most frequent; occasionally compact patches or streaks of specialized hair and short plume scales occur. Eyes variable. Antennae with nudum confined to club. Palpi variable. Male genitalia with uncus undivided or (more commonly) divided into separated lobes; brachia usually present, but never bifurcate; juxta usually furca-like, but occasionally reduced to a small plate or otherwise modified; saccus usually not or only very slightly developed.
<==Polyommatini
|--Zizula Chapman 1910WF02
| `--Z. hylax (Fabricius 1775)B2000 [=Papilio hylaxWF02; incl. Lycaena gaika Trimen 1862WF02, *Zizula gaikaWF02]
| |--Z. h. hylaxB12
| `--Z. h. attenuata (Lucas 1890)B12
|--Famegana Eliot 1973WF02
| `--*F. alsulus (Herrich-Schäffer 1869)WF02 (see below for synonymy)
| |--F. a. alsulusWF02
| `--F. a. taiwana (Sonan 1938) [=Ziazera alsulus taiwana, Zizina alsulus taiwana]WF02
|--Lampides Hübner 1819 [=Cosmolyce Toxopeus 1927, Lampidella Hemming 1933]WF02
| |--*L. boeticus (Linnaeus 1767)WF02 (see below for synonymy)
| |--L. agricolaO90
| |--L. alsulusO90
| `--L. conferendaB1900b
|--Petrelaea Toxopeus 1929WF02
| |--P. dana (de Nicéville 1884) [=Nacaduba dana]WF02
| | |--P. d. danaWF02
| | `--*P. d. varia Toxopeus 1929WF02
| `--P. tombugensis (Rober 1886)B2000
|--Euchrysops Butler 1900WF02
| |--*E. cnejus (Fabricius 1798) [=Hesperia cnejus, Catochrysops cnejus]WF02
| | |--E. c. cnejusB12
| | `--E. c. cnidus Waterhouse & Lyell 1914B12
| `--E. cyclopteris [=Catochrysops cyclopteris]B01c
|--Castalius sectionWF02
| |--Castalius Hübner 1819WF02
| | |--*C. rosimon (Fabricius 1775) [=Papilio rosimon]WF02
| | `--C. melaenaB01a
| `--Tarucus Moore 1881WF02
| |--*T. theophrastus (Fabricius 1793) [=Hesperia theophrastus]WF02
| `--T. telicanusB01a
|--Una sectionWF02
| |--Una de Nicéville 1890B2000
| | `--*U. usta (Distant 1886) [=Zizera usta]WF02
| `--Orthomiella de Nicéville 1890WF02
| |--*O. pontis (Elwes 1887) [=Chilades pontis, Una pontis]WF02
| |--O. fukienensis Forster 1941 [=O. pontis fukienensis]WF02
| |--O. rantaizana Wileman 1910 [=O. pontis rantaizana, Una rantaizana; incl. O. lucida Forster 1942]WF02
| `--O. sinensis (Elwes 1887) [=Chilades sinensis, O. pontis var. sinensis, Una pontis sinensis]WF02
|--Nacaduba sectionWF02
| |--NacadubaB2000
| |--ProsotasWF02
| `--Ionolyce Toxopeus 1929WF02
| `--I. helicon (Felder 1860) [=Lycaena helicon]WF02
| |--I. h. heliconB2000
| |--I. h. caracallaB2000
| |--I. h. hyllus (Waterhouse & Lyell 1914)B2000
| |--*I. h. javanica Toxopeus 1929WF02
| `--I. h. viola (Moore 1877) [=Lampides viola, Nacaduba viola]WF02
|--Pithecops Horsfield 1828 [incl. Eupsychellus Röber 1891]WF02
| |--*P. hylax Horsfield 1828WF02
| |--P. corvus Fruhstorfer 1919WF02
| | |--P. c. corvusWF02
| | `--P. c. correctus Cowan 1966WF02
| |--P. dionisius (Boisduval 1832)B2000 [=Lycaena dionisiusWF02, *Eupsychellus dionisiusWF02]
| |--P. fulgens Doherty 1889 [=Pithecopis (l. c.) fulgens]WF02
| | |--P. f. fulgensI92
| | |--P. f. tsushimanus Shirôzu & Urata 1957I92
| | `--P. f. urai Bethune-Baker 1913 [=P. nihana urai]WF02
| `--P. ryukyuensis Shirôzu 1964WF02 [=P. fulgens ryukyuensisWF02, P. corvus ryukyuensisI92]
|--Upolampes sectionWF02
| |--Discolampa Toxopeus 1929 [=Ethion Shirôzu & Saigusa 1962]WF02
| | `--*D. ethion (Westwood 1851) [=Lycaena ethion, Castalius ethion, *Ethion ethion]WF02
| |--Syntarucus Butler 1901WF02
| | |--‘Papilio’ pirithous L. 1767 [incl. P. telicanus Lang 1789, *Syntarucus telicanus]WF02
| | `--S. plinius (Fabricius 1793) [=Hesperia plinius, Leptotes plinius, Tarucus plinius]WF02
| | |--S. p. pliniusB2000
| | `--‘Leptotes’ p. pseudocassius (Murray 1873)B2000
| `--Caleta Fruhstorfer 1922 [incl. Pycnophallium Toxopeus 1929]WF02
| |--*C. caleta (Hewitson 1876) [=Lycaena caleta]WF02
| |--C. decidia (Hewitson 1876) [=Lycaena decidia, Caleta caleta decidia]WF02
| |--C. elna (Hewitson 1876) [=Lycaena elna, Castalius elna]WF02
| | |--C. e. elnaWF02
| | `--C. e. noliteia (Fruhstorfer 1918) [=Castalius elna noliteia]WF02
| `--C. roxus (Godart 1824) [=Polyommatus roxus, *Pycnophallium roxus]WF02
| |--C. r. roxusWF02
| `--C. r. roxana (de Nicéville 1897) [=Castalius roxus roxana]WF02
|--Catochrysops Boisduval 1832WF02, B2000
| |--*C. strabo (Fabricius 1793) [=Hesperia strabo]WF02
| |--C. amasea Waterhouse & Lyell 1914B2000
| |--C. asopusB01
| |--C. carsoni Butler 1901B01c
| |--C. celaeusB01c
| |--C. hypoleucus [incl. C. gigantea]B01a
| |--C. negusB01d
| |--C. osirisB01a
| |--C. panormus (Felder 1860) [=Lycaena panormus]WF02
| | |--C. p. panormusB12
| | |--C. p. exiguus (Distant 1886) [=Everes panormus exiguus]WF02
| | |--C. p. papuana Tite 1959B2000
| | `--C. p. platissa (Herrich-Schäffer 1869)B12
| |--C. patriciaB01c
| |--C. peculiarisB01a
| |--C. phasma Butler 1901B01c
| `--C. taitensisPP02
|--Zizeeria sectionWF02
| |--Pseudozizeeria Beuret 1955WF02
| | `--*P. maha (Kollar 1844) [=Lycaena maha, Zizeeria maha, Zizera maha]WF02
| | |--P. m. mahaTYM08
| | |--‘Zizeeria’ m. kashikurai Nakamura 1930TYM08
| | `--P. m. okinawana (Matsumura 1929) [=Zizera maha okinawana, Zizeeria maha okinawana]WF02
| |--Zizeeria Chapman 1910B2000
| | |--*Z. karsandra (Moore 1865) [=Polyommatus lysimon karsandra, Zizera lysimon karsandra]WF02
| | |--Z. knysna (Trimen 1862) [=Lycaena knysna]WF02
| | `--Z. otisC70
| | |--Z. o. otisC70
| | `--Z. o. labradusC70
| `--Zizina Chapman 1910WF02
| |--Z. otis (Fabricius 1787) [=Papilio otis, Zizera otis]WF02
| | |--Z. o. otisB12
| | |--Z. ‘labradus’ labdalon Waterhouse & Lyell 1914B2000
| | |--*Z. o. labradus (Godart 1824)WF02, B12 [=Polyommatus labradusWF02; incl. Z. otis mangoensisPP02]
| | `--Z. o. riukuensis (Matsumura 1929) [=Zizera otis riukuensis]WF02
| `--Z. oxleyiPPE03
|--Everes sectionWF02
| |--EveresB2000
| |--TongeiaWF02
| |--Sinocupido Lee 1963WF02
| | `--*S. lokiangensis Lee 1963WF02
| |--Shijimia Matsumura 1919WF02
| | `--*S. moorei (Leech 1889) [=Lycaena moorei]WF02
| | |--S. m. mooreiWF02
| | `--S. m. taiwana Matsumura 1919WF02
| |--Bothrinia Chapman 1909WF02
| | |--*B. chennellii (de Nicéville 1884) [=Cyaniris chennellii]WF02
| | `--B. nebulosa (Leech 1890) [=Cyaniris nebulosa, Lycenopsis nebulosa]WF02
| | |--B. n. nebulosaWF02
| | `--B. n. leechi Forster 1941WF02
| `--Cupido Schrank 1801WF02
| |--*C. minimus (Fuessly 1775) [=Papilio minimus]WF02
| | |--C. m. minimusWF02
| | `--C. m. qilianus Murayama 1983WF02
| |--C. buddhista (Alphéraky 1881) [=Lycaena buddhista, Zizera lorquinii buddhista]WF02
| `--C. prosecusa (Erschoff 1874) [=Lycaena prosecusa, Zizera prosecusa]WF02
|--Jamides Hübner 1819WF02, B2000
| |--*J. bochus (Stoll 1782) [=Papilio bochus]WF02
| | |--J. b. bochusPP02
| | |--J. b. candrenaPP02
| | `--J. b. formosanus Fruhstorfer 1909WF02
| |--J. alecto (Felder 1860) [=Cupido alecto]WF02
| | |--J. a. alectoWF02
| | |--J. a. alocina Swinhoe 1915WF02
| | `--J. a. dromicus (Fruhstorfer 1910) [=Lampides alecto dromicus, L. elpis dromicus]WF02
| |--J. aleuas (Felder & Felder 1865)B2000
| | |--J. a. aleuasB2000
| | `--J. a. coelestis (Miskin 1891)B2000
| |--J. amarauge Druce 1891B2000
| |--J. celeno (Cramer 1775) [=Papilio celeno, Lampides celeno]WF02
| | |--J. c. celenoS72
| | `--J. c. aelianusS72
| |--J. cyta (Boisduval 1832)B2000
| | |--J. c. cytaB2000
| | `--J. c. claudia (Waterhouse & Lyell 1914)B2000
| |--J. elpis (Godart 1824) [=Polyommatus elpis]WF02
| |--J. nemophila (Butler 1876)B2000
| |--J. phaseli (Mathew 1889)B12
| `--J. pura (Moore 1886) [=Lampides pura, L. celeno pura]WF02
|--Polyommatus sectionWF02
| |--PolyommatusBH07
| |--AlbulinaWF02
| |--AgriadesWF02
| |--LycaeidesWF02
| |--AgrodiaetusWF02
| |--CyanirisWF02
| |--PlebejusWF02
| |--Eumedonia Forster 1938WF02
| | `--*E. eumedon (Esper 1780) [=Papilio eumedon]WF02
| | |--E. e. eumedonWF02
| | |--E. e. antiqua Staudinger 1899WF02
| | `--E. e. privata (Staudinger 1895) [=Lycaena eumedon privata]WF02
| |--Freyeria Courvoisier 1920WF02
| | |--*F. trochylus (Freyer 1845)WF02 [=Lycaena trochylusWF02, Chilades trochilus (l. c.)WF02, Plebeius trochilusB1900a]
| | `--F. putli (Kollar 1844) [=Lycaena putli, Chilades trochilus (l. c.) putli, Freyeria trochylus putli]B12
| | |--F. p. putliWF02
| | `--F. p. formosana Matsumura 1919 [=F. trochylus formosana, Chilades trochylus formosanus]WF02
| |--Vacciniina Tutt 1909WF02
| | `--*V. optilete (Knoch 1781) [=Papilio optilete]WF02
| | |--V. o. optileteI92
| | |--V. o. daisetsuzana (Matsumura 1926)I92
| | `--V. o. sibirica (Staudinger 1892) [=Lycaena sibirica]WF02
| |--Aricia R. L. 1817WF02
| | |--*A. agestis (Denis & Schiffermüller 1775) [=Papilio agestis]WF02
| | |--A. allous (Geyer 1836) [=Papilio artaxerxes allous]WF02
| | |--A. artaxerxes (Fabricius 1781)BH07
| | |--A. chinensis (Murray 1874) (see below for synonymy)WF02
| | |--A. eumedon (Esper 1782)BH07
| | | |--A. e. eumedonF04
| | | `--A. e. axarquia Gil-T. 2004F04
| | `--A. mandschurica (Staudinger 1892)WF02 [=Lycaena mandschuricaWF02, L. astrarche var. mandschuricaB89]
| `--Chilades Moore 1881WF02
| |--*C. lajus (Stoll 1780) [=Papilio lajus, C. laius (l. c.)]WF02
| | |--C. l. lajusWF02
| | `--C. l. koshunensis Matsumura 1919WF02
| |--C. alberta Butler 1901B01c
| |--C. pandava (Horsfield 1829) [=Lycaena pandava, Catochrysops pandava, Euchrysops pandava]WF02
| | |--C. p. pandavaWF02
| | `--C. p. peipatria Hsu 1989WF02
| `--C. yunnanensis Watkins 1927WF02
|--Lycaenopsis sectionWF02
| |--CelastrinaWF02
| |--Lestranicus Eliot & Kawazoé 1983WF02
| | `--*L. transpectus (Moore 1879)WF02 (see below for synonymy)
| |--Celatoxia Eliot & Kawazoé 1983WF02
| | |--*C. albidisca (Moore 1884) [=Cyaniris albidisca]WF02
| | `--C. marginata (de Nicéville 1884) [=Cyaniris marginata, Lycaenopsis marginata]WF02
| |--Monodontides Toxopeus 1927WF02
| | |--*M. argioloides (Rothschild 19160 [=Nacaduba argioloides]WF02
| | `--M. musina (Snellen 1892) [=Lycaena musina]WF02
| |--Udara Toxopeus 1928WF02
| | |--*U. dilecta (Moore 1879) (see below for synonymy)WF02
| | |--U. albocaerulea (Moore 1879) (see below for synonymy)WF02
| | `--U. tenella (Miskin 1891)B2000
| |--Callenya Eliot & Kawazoé 1983WF02
| | `--*C. melaena (Doherty 1889) [=Cyaniris melaena]WF02
| | |--C. m. melaenaWF02
| | `--C. m. shonen (Esaki 1932) (see below for synonymy)WF02
| |--Acytolepis Toxopeus 1927WF02
| | `--*A. puspa (Horsfield 1828)WF02 (see below for synonymy)
| | |--A. p. puspaWF02
| | |--A. p. gisca (Fruhstorfer 1910) (see below for synonymy)WF02
| | `--A. p. myla (Fruhstorfer 1909) (see below for synonymy)WF02
| |--LycaenopsisB2000
| | |--L. argiolusC37
| | | |--L. a. argiolusC37
| | | `--L. a. pseudargiolusC37
| | |--‘Cyaniris’ catreus De Nicéville 1895B1900b
| | `--‘Cupido’ deliciosa Pagenstecher 1896 [=Cyaniris deliciosa]B1900b
| |--Megisba Moore 1881 [incl. Pathalia Moore 1884]WF02
| | |--M. malaya (Horsfield 1828) [=Lycaena malaya; incl. *M. thwaitesi Moore 1881]WF02
| | | |--M. m. malayaWF02
| | | `--M. m. sikkima Moore 1884 [incl. M. malaya volubilis Fruhstorfer 1918]WF02
| | |--‘*Pathalia’ albidisca Moore 1884WF02
| | `--M. strongyle (Felder 1860)B2000
| | |--M. s. strongyleB2000
| | `--M. s. nigra (Miskin 1890)B2000
| `--Neopithecops Distant 1884 [incl. Papua Röber 1892 non Ragonot 1889, Parapithecops Moore 1884]WF02
| |--N. zelmora (Butler 1870) [=Pithecops zalmora]WF02
| | |--N. z. zelmora (see below for synonymy)WF02
| | `--*N. z. dharma (Moore 1881) [=Pithecops dharma]WF02
| |--‘*Parapithecops’ gaura Moore 1884WF02
| |--N. lucifer (Röber 1886)B2000 [=Plebeius luciferWF02, *Papua luciferWF02]
| | |--N. l. luciferB2000
| | `--N. l. heria (Fruhstorfer 1919)B2000
| `--‘Papua’ rhabdotaP27
`--Glaucopsyche sectionWF02
| i. s.: Palaeophilotes Forster 1938WF02
| `--*P. triphysina (Staudinger 1892) [=Lycaena triphysina]WF02
| |--P. t. triphysinaWF02
| `--P. t. yuliana Lee 1962WF02
| Shijimiaeoides Beuret 1958WF02
| |--S. divina (Fixen 1887) [=Lycaena divina, Sinia divina]WF02
| | |--S. d. divinaI92
| | `--*S. d. barine (Leech 1893)WF02, I92 [=Lycaena barineWF02]
| |--S. battoidesUSDI77
| | |--S. b. battoidesUSDI77
| | `--S. b. allyniUSDI77
| `--S. enoptesUSDI77
| |--S. e. enoptesUSDI77
| `--S. e. smithiUSDI77
| Subsulanoides Koiwaya 1989WF02
| `--*S. nagata Koiwaya 1989WF02
|--+--+--Otnjukovia tatjanaAV04
| | `--Turanana cytisAV04
| `--Iolana Bethune-Baker 1914AV04, WF02
| |--*I. iolas (Ochsenheimer 1816) [=Lycaena iolas]WF02
| `--I. gigantea (Grum-Grshimaïlo 1885) [=Lycaena gigantea]WF02
| |--I. g. giganteaWF02
| `--I. g. gilgitica (Tytler 1926) [=Polyommatus gigantea gilgitica, Lycaena gigantea gilgitica]WF02
|--Phengaris Doherty 1891AV04
| |--P. daitozana Wileman 1908AV04, WF02 [=Lycaena atroguttata daitozanaWF02, P. dastozana (l. c.)WF02]
| `--+--MaculineaAV04
| `--+--*P. atroguttata (Oberthür 1876) [=Lycaena atroguttata]WF02
| | |--P. a. atroguttataWF02
| | |--P. a. formosana (Matsumura 1926) [=Lycaena atroguttata formosana]WF02
| | `--P. a. juenana (Forster 1940) [=Glaucopsyche atroguttata juenana]WF02
| `--P. abida Leech 1893AV04, WF02 [=P. atroguttata var. abidaWF02, Lycaena atroguttata abidaWF02]
`--+--+--PseudophilotesAV04
| | |--P. baviusAV04
| | `--+--P. batonAV04
| | `--P. vicramaAV04
| `--Scolitantides Hübner 1819AV04, WF02 [Scolitantiditi]
| `--*S. orion (Pallas 1771) [=Papilio orion]WF02
| |--S. o. orionWF02
| |--S. o. coreana (Matsumura 1926) [=Lycaena orion coreana, S. orion matsumuranus Bryk 1946]WF02
| `--S. o. ornata (Staudinger 1892) [=Lycaena ornata]WF02
`--+--GlaucopsycheAV04
`--Caerulea Forster 1938AV04, WF02
|--*C. coelestis (Alphéraky 1897) [=Lycaena coeligena var. coelestis, Maculinea coelestis]WF02
| |--C. c. coelestis [incl. Lycaena coeli Oberthür 1908]WF02
| |--C. c. dubernardi Hemming 1931WF02
| `--C. c. sora Sugiyama 1999WF02
`--C. coeligena (Oberthür 1876) [=Lycaena coeligena, Maculinea coeligena]WF02
|--C. c. coeligenaWF02
`--C. c. pratti Hemming 1931WF02
Polyommatini incertae sedis:
Catopyrops Toxopeus 1929B2000
|--C. ancyra (Felder 1860)B2000
| |--C. a. ancyraB2000
| |--C. a. exponensB2000
| `--C. a. mysia (Waterhouse & Lyell 1914)B2000
`--C. florinda (Butler 1877)B2000
|--C. f. florindaB12
|--C. f. estrella (Waterhouse & Lyell 1914)B12
`--C. f. halys Waterhouse 1934B2000
LepidochrysopsAV04
PhaedrotesAV04
ApellesAV04
PhilotesAV04
|--P. battoidesS72
| |--P. b. battoidesS72
| `--P. b. bernardinoS72
|--P. enoptesS72
`--P. sonorensisL72
PraephilotesAV04
PhilotiellaAV04
MicropsycheAV04
RubrapterusAV04
BajluanaAV04
LachidesAV04
Erysichton Fruhstorfer 1916B2000
|--E. lineata (Murray 1874)B2000
`--E. palmyra (Felder 1860)B2000
|--E. p. palmyraB2000
`--E. p. tasmanicus (Miskin 1890)B2000
Danis Fabricius 1807B2000
`--D. danis (Cramer 1775)B2000
|--D. d. danisB2000
|--D. d. serapis Miskin 1891B2000
`--D. d. syrius Miskin 1890B2000
Psychonotis Toxopeus 1930B2000
|--P. caelius (Felder 1860)B2000
| |--P. c. caeliusB2000
| `--P. c. taygetus (Felder & Felder 1865) (see below for synonymy)B2000
`--P. hymetus (Felder 1860) [=Danis hymetus]B2000
Perpheres Hirowatari 1992B2000
TheclinesthesB2000
Sahulana Hirowatari 1992B2000
`--S. scintillata (Lucas 1889) [=Theclinesthes scintillata]B2000
Neolucia Waterhouse & Turner 1905B2000
|--N. agricola (Westwood 1851)B2000
| |--N. a. agricolaB2000
| |--N. a. insulana Waterhouse & Lyell 1914B2000
| `--N. a. occidens Waterhouse & Lyell 1914B2000
|--N. hobartensis (Miskin 1890)B2000
| |--N. h. hobartensisB2000
| `--N. h. monticola Waterhouse & Lyell 1914B2000
`--N. mathewi (Miskin 1890)B2000
Leptotes Scudder 1876B2000
|--L. cassius (Cramer 1775)FS54
|--L. marinaS72
`--L. trigemmatusH11
Cyclirius Butler 1897B2000
EuphilotesAV04
|--E. battoidesAV04
`--E. enoptesAV04
UranothaumaAJV-W99
|--U. cordatusB01b
`--U. nubiferB01b
*Acytolepis puspa (Horsfield 1828)WF02 [=Polyommatus puspaWF02, Cyaniris puspaB1900b; incl. C. crissa De Nicéville 1895B1900b, C. puspa var. lilacea Hampson 1889B1900b]
Acytolepis puspa gisca (Fruhstorfer 1910) [=Cyaniris paspa gisca, Lycaenopsis paspa gisca; incl. Celastrina puspa barneyi Corbet 1941, Cyaniris paspa hermagoras Fruhstorfer 1910]WF02
Acytolepis puspa myla (Fruhstorfer 1909) [=Cyaniris puspa myla, Celastrina puspa myla, Lycaenopsis puspa myla; incl. Ce. puspa formosana Bethune-Baker 1914, Cyaniris postimacula Matsumura 1929]WF02
Aricia chinensis (Murray 1874) [=Lycaena chinensis non Chrysophanus phlaeas chinensis Felder 1862 (not preoc. if in dif. gen.)]WF02
Callenya melaena shonen (Esaki 1932) [=Lycaenopsis transpectus shonen, Celastrina minima shonen, Cyaniris latimargo Matsumura 1929 non Moore 1883]WF02
*Famegana alsulus (Herrich-Schäffer 1869)WF02 [=Lycaena alsulusWF02, Zizeeria alsulusB2000, Zizera labradus alsulusWF02, Zizina alsulusB2000]
*Lampides boeticus (Linnaeus 1767)WF02 [=Papilio boeticusWF02, *Cosmolyce boeticusWF02, *Lampidella boeticusWF02, Polyommatus boeticusB01b, Po. baeticus (l. c.)WF02]
*Lestranicus transpectus (Moore 1879)WF02 [=Polyommatus transpectusWF02, Cyaniris transpectusB1900b, Lycaenopsis transpectusWF02; incl. C. latimargo Moore 1883B1900b]
Neopithecops zelmora zelmora (Butler 1870) [incl. Pithecops zalmora dolona Fruhstorfer 1919, P. zalmora fedora Fruhstorfer 1919, P. nihina Moore 1878]WF02
Psychonotis caelius taygetus (Felder & Felder 1865) [incl. P. caelius salamandri (Macleay 1866), P. caelius taletum (Waterhouse & Lyell 1914)]B2000
Udara albocaerulea (Moore 1879) [=Polyommatus albocaeruleus, Celastrina albocaerulea, Cyaniris albocaeruleus, Lycaenopsis albocaeruleus; incl. Cy. ceyx amadis Fruhstorfer 1910, Celastrina albocaerulea amandis, Lycaenopsis albocaeruleus sauteri Fruhstorfer 1917, Celastrina albocaerulea sauteri]WF02
*Udara dilecta (Moore 1879) [=Polyommatus dilectus, Celastrina dilectus, Cyaniris dilectus, Lycaenopsis cardia dilectus; incl. L. cardia dilectina Fruhstorfer 1917, Cyaniris dilectus hainana Fruhstorfer 1910, Cy. dilectus hermonthis Fruhstorfer 1909, Celastrina dilecta hermonthis]WF02
*Type species of generic name indicated
References
[AJV-W99] Ackery, P. R., R. de Jong & R. I. Vane-Wright. 1999. The butterflies: Hedyloidea, Hesperioidea and Papilionoidea. In: Kristensen, N. P. (ed.). Handbuch der Zoologie vol. 4. Arthropoda: Insecta pt 35. Lepidoptera, Moths and Butterflies vol. 1. Evolution, Systematics, and Biogeography pp. 263–300. Walter de Gruyter.
[AV04] Als, T. D., R. Vila, N. P. Kandul, D. R. Nash, S.-H. Yen, Y.-F. Hsu, A. A. Mignault, J. J. Boomsma & N. E. Pierce. 2004. The evolution of alternative parasitic life histories in large blue butterflies. Nature 432: 386–390.
[BH07] Bálint, Z., Z. E. Horváth, K. Kertész, Z. Vértesy & L. P. Biró. 2007. Observations on scale structures and spectroscopic properties of Polyommatus lycaenid butterflies (Lepidoptera: Lycaenidae). Annales Historico-Naturales Musei Nationalis Hungarici 99: 115–127.
[B89] Blachier, C. 1889. Une Boarmia nouvelle, variétés inédites de lépidoptères, et chenille d’une hémérophile. Annales de la Société Entomologique de France, 6e série 9: 255–260, pl. 4.
[B2000] Braby, M. F. 2000. Butterflies of Australia: their identification, biology and distribution vol. 2. CSIRO Publishing: Collingwood (Victoria).
[B12] Braby, M. F. 2012. The butterflies of El Questro Wilderness Park, with remarks on the taxonomy of the Kimberley fauna, Australia. Records of the Western Australian Museum 27 (2): 161–175.
[B1900a] Butler, A. G. 1900a. On a second collection of butterflies obtained by Mr. Edward M. de Jersey in Nyasaland. Annals and Magazine of Natural History, series 7, 5: 59–63.
[B1900b] Butler, A. G. 1900b. A list of the species of Cyaniris, a well-known group of the family Lycaenidae. Annals and Magazine of Natural History, series 7, 5: 441–451.
[B01a] Butler, A. G. 1901a. An account of a collection of butterflies made by the Rev. K. St. Aubyn Rogers between Mombasa and the forests of Taveta. Annals and Magazine of Natural History, series 7, 7: 22–35.
[B01b] Butler, A. G. 1901b. An account of a collection of butterflies obtained by Lord Delamere, chiefly at Munisu, near Mount Kenya. Annals and Magazine of Natural History, series 7, 7: 197–203.
[B01c] Butler, A. G. 1901c. Descriptions of new species of Lycaenidae in the collection of the British Museum. Annals and Magazine of Natural History, series 7, 7: 288–291.
[B01d] Butler, A. G. 1901d. On a collection of butterflies from the Uganda Protectorate, forwarded by C. Steuart Betton, Esq., in 1900. Annals and Magazine of Natural History, series 7, 7: 562.
[C37] Clark, A. H. 1937. The butterflies of Virginia. Proceedings of the Entomological Society of Washington 39 (5): 116.
[C70] Common, I. F. B. 1970. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 765–866. Melbourne University Press.
[F04] Fernández, J. 2004. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (VIII). Graellsia 60 (2): 217–243.
[FS54] Franz, E., & H. Schröder. 1954. Tagfalter (Lep. Rhopalocera) aus El Salvador. Senckenbergiana Biologica 35: 75–87.
[H11] Hilty, S. L. 2011. Family Thraupidae (tanagers). In: Hoyo, J. del, A. Elliott & D. A. Christie (eds) Handbook of the Birds of the World vol. 16. Tanagers to New World Blackbirds pp. 46–329. Lynx Edicions: Barcelona.
[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.
[L72] Langston, R. L. 1972. The Sonora blue—1971 the earliest season for the north. Pan-Pacific Entomologist 48 (1): 67.
[O90] Olliff, A. S. 1890. On Rhopalocera from Mt. Kosciusko, New South Wales. Proceedings of the Linnean Society of New South Wales, series 2, 4 (3): 619–624.
[PP02] Patrick, B., & H. Patrick. 2002. Butterflies of Fiji. Weta 24: 5–12.
[PPE03] Patrick, B., H. Patrick & E. Edwards. 2003. Tiger moths and pheromones. Weta 25: 14–17.
[P27] Philpott, A. 1927. The maxillae in the Lepidoptera. Transactions and Proceedings of the New Zealand Institute 57: 721–746.
[S72] Shields, O. 1972. Flower visitation records for butterflies (Lepidoptera). Pan-Pacific Entomologist 48 (3): 189–203.
[TYM08] Tennent, W. J., M. Yasuda & K. Morimoto. 2008. Lansania Journal of arachnology and zoology—a rare and obscure Japanese natural history journal. Archives of Natural History 35 (2): 252–280.
[USDI77] United States Department of the Interior. 1977. Endangered and threatened wildlife and plants—republication of list of species. Federal Register 42: 36420–36431.
[WF02] Wang M. & Fan X. 2002. Butterflies Fauna Sinica: Lycaenidae. Henan Science and Technology Publishing House.