Belongs within: Dicranideae.
Contains: Trichostomum, Tortella, Bryoerythrophylleae, Leptodontieae, Barbula, Didymodon, Weissia, Hyophileae, Tortula, Acaulon, Phascum, Pottia, Syntrichia.
The Pottiaceae is a large, diverse family of mosses found worldwide, divided between about eighty genera. Members of the family are commonly small and found in harsh environments such as rocks and disturbed soil, primarily in calcareous habitats. Members of the tribe Trichostomeae usually have lanceolate, tapering leaves whereas members of the Pottieae more commonly have broad leaves. The genus Cinclidotus has large, solid, bordered leaves and is found in aquatic or riparian habitats (Dixon 1924).
Characters (from Smith 2004): Plants usually small, often xeromorphic. Leaves linear to spathulate or ovate, acuminate to rounded; margins entire to dentate, usually unbordered; costa usually strong, in section with one or two stereid bands; basal cells rectangular, more or less hyaline, cells above hexagonal or quadrate, often papillose, sometimes obscure. Setae very short to long, usually straight; capsules erect or inclined, globose to cylindrical, straight or curved, cleistocarpous or dehiscent; peristome absent, rudimentary, imperfect or perfect, teeth straight or spirally coiled, entire, perforated or variously cleft or divided into filiform segments.
<==Pottiaceae [Tortulaceae] |--Trichostomoideae [Trichostomeae]S04 | |--TrichostomumSK02 | |--TortellaS04 | |--Eucladium Bruch & Schimp. in Bruch, Schimp. & Gümbel 1846S04 | | `--E. verticillatum (Brid.) Bruch & Schimp. in Bruch, Schimp. & Gümbel 1846S04 (see below for synonymy) | |--Pottiopsis Blockeel & Sm. 1998S04 | | `--P. caespitosa (Bruch ex Brid.) Blockeel & Sm. 1998S04 (see below for synonymy) | |--Paraleptodontium Long 1982S04 | | `--P. recurvifolium (Taylor) Long 1982S04 (see below for synonymy) | `--Pleurochaete Lindb. 1864S04 | |--P. luteolaJ87 | `--P. squarrosa (Brid.) Lindb. 1864S04 [=Barbula squarrosaD24, Tortella squarrosaOZB-O03] |--MerceyoideaeS04 | |--BryoerythrophylleaeS04 | |--LeptodontieaeS04 | `--BarbuleaeS04 | |--BarbulaD24 | |--DidymodonS04 | |--Scopelophila (Mitt.) Lindb. 1872S04 | | `--S. cataractae (Mitt.) Broth. 1902S04 | |--Gyroweisia Schimp. 1876 (nom. cons.)S04 | | |--G. reflexa (Brid.) Schimp. 1876 [=Weissia reflexa]S04 | | `--G. tenuis (Schrad. ex Hedw.) Schimp. 1876S04 (see below for synonymy) | |--Molendoa Lindb. 1878S04 | | |--M. hornschuchiana (Hook.) Lindb. ex Limpr. 1886 (see below for synonymy)OZB-O03 | | |--M. sendtneriana (Bruch & Schimp.) Limpr. 1886 [=M. hornschuchiana ssp. sendtneriana]OZB-O03 | | `--M. warburgii (Crundw. & Hill) Zander 1993 [=Anoectangium warburgii]S04 | |--Gymnostomum Nees & Hornsch. in Nees et al. 1823S04 | | |--G. aeruginosum Smith 1804SK02 (see below for synonymy) | | |--G. calcareum Nees & Hornsch. in Nees et al. 1823S04 (see below for synonymy) | | |--G. laxireteB-OO10 | | `--G. viridulum Brid. 1826S04 (see below for synonymy) | `--Anoectangium Schwägr. 1811 (nom. cons.)S04 | |--A. aestivum (Hedw.) Mitt. 1869SK02 (see below for synonymy) | | |--A. a. var. aestivumD24 | | `--A. ‘compactum’ var. pellucidum [incl. A. marinum]D24 | |--A. bellii Broth. ex Dixon 1926SK02 | `--A. thomsoniiN02 `--PottioideaeS04 | i. s.: WeissiaOZB-O03 | Hilpertia Zander 1989OZB-O03 | `--H. velenovskyi (Schiffn.) Zander 1989 [=Tortula velenovskyi]OZB-O03 |--HyophileaeS04 `--PottieaeD24 |--TortulaSK02 |--AcaulonD24 |--PhascumS04 |--PottiaS04 |--SyntrichiaS04 |--Leptobarbula Schimp. 1876S04 | `--L. berica (De Not.) Schimp. 1876S04 |--Leptophascum (Müll. Hal.) Guerra & Cano 2000S04 | `--*L. leptophyllum (Müll. Hal.) Guerra & Cano 2000SK02, S04 (see below for synonymy) |--Hennediella Paris 1896S04 | |--H. heimii (Hedw.) Zander 1993SK02 (see below for synonymy) | |--H. macrophylla (Br.) Paris 1896 [incl. Tortula brevis]S04 | `--H. stanfordensis (Steere) Blockeel 1990S04 (see below for synonymy) |--Protobryum (Limpr.) Guerra & Cano 2000S04 [=Mildeella Limpricht 1885 (preoc.)OZB-O03] | `--P. bryoides (Dicks.) Guerra & Cano 2000S04 (see below for synonymy) | |--P. b. var. bryoidesD24 | |--‘Pottia’ b. var. piliferaD24 | `--‘Pottia’ b. var. thornhilliiD24 `--Microbryum Schimp. 1860S04 |--M. brevicaule (Taylor) Zander 1993 (see below for synonymy)SK02 |--M. curvicolle (Hedw.) Zander 1993 [=Phascum curvicolle]S04 |--M. floerkeanum (Weber & Mohr) Schimp. 1860S04 (see below for synonymy) `--M. rectum (With.) Zander 1993S04 [=Phascum rectumD24, Pottia rectaS04] Pottiaceae incertae sedis: Timmiella barbuloidesFHH01 Phasconica Müll.Hal. 1882SK02 Semibarbula orientalis (Weber) Wijk & Margad. 1959 [=Trichostomum orientale Weber 1804]SG07 Calymperastrum Stone 1986SK02 `--C. latifolium (Hampe) Stone 1986 [=Calymperes latifolium]SK02 Chenia Zander 1989 non Sheng 1963 (ICZN)SK02 Gymnostomiella Fleischer 1904SK02 `--G. vernicosa (Hooker) Fleischer 1904 [=Gymnostomum vernicosum Hooker 1836]B05 Phascopsis Stone 1980SK02 `--P. rubicunda Stone 1980SK02 Stonea Zander 1989SK02 `--S. oleaginosa (Stone) Zander 1989 [=Tortula oleaginosa Stone 1978]SK02 Pseudosymblepharis Broth. 1924SK02 |--P. bombayensis (Müll.Hal.) Sollman 1996 [incl. P. angustata (Mitt.) Hilp. 1933]SK02 |--P. schimperianusJ87 `--P. subduriuscula (Müll.Hal.) Chen 1941SK02 Crossidium Jur. 1882SK02 |--C. davidai Catches. 1980SK02 `--C. geheebii (Broth.) Broth. 1902 [=Barbula geheebii (nom. inv.)]SK02 Splachnobryum Müll.Hal. 1869SK02 |--S. obtusum (Brid.) Müll.Hal. 1869 (see below for synonymy)SK02 `--S. wiemansii Fleischer 1904SK02 Uleobryum Broth. 1906SK02 |--U. occultum (Roth) Zander 1993 [incl. U. curtisii Stone 1984]SK02 `--U. peruvianum Broth. 1906SK02 Trachycarpidium Broth. 1901SK02 |--T. brisbanicum (Müll.Hal.) Stone 1975 (see below for synonymy)SK02 `--T. verrucosumSK02 HydrogoniumSG07 |--H. aneitense [=Barbula (Hydrogonium) aneitensis Brotherus & Watts 1915]BW15 |--H. arcuatum (Griffith) Wijk. & Marg. 1958 [=Barbula arcuata Griffith 1842]B05 `--H. consanguineum (Thwaites & Mitt.) Hilp. 1933 [=Tortula consanguinea Thwaites & Mitt. 1873]SG07 Calyptopogon (Mitt.) Broth. 1902SK02 `--C. mnioides (Schwägr.) Broth. 1902 (see below for synonymy)SK02 |--C. m. var. mnioidesSK02 `--‘Streptopogon’ m. var. prostratus [=Tortula prostrata]SK02 Goniomitrium Hooker & Wilson 1846SK02 `--G. acuminatum Hooker & Wilson 1846SK02 |--G. a. ssp. acuminatum [incl. Physcomitrium goniomitrium]SK02 `--G. a. ssp. enerve (Hooker & Wilson) Fife & Seppelt 2001 (see below for synonymy)SK02 Triquetrella Müll.Hal. 1897SK02 |--T. fragilis Müll.Hal. 1897SK02 |--T. papillata (Hooker & Wilson) Broth. 1902 (see below for synonymy)SK02 |--T. richardsiae Müll.Hal. 1897SK02 `--T. tasmanica (Broth.) Granzow 1989 (see below for synonymy)SK02 Tetrapterum Hampe ex Jaeger 1869SK02 |--T. brachypelma (Müll.Hal.) Broth. 1924 (see below for synonymy)SK02 |--T. cylindricum (Taylor) Jaeger 1869 (see below for synonymy)SK02 |--T. sullivanii (Müll.Hal.) Broth. 1924 (see below for synonymy)SK02 |--T. tetrapteroides (Müll.Hal.) Broth. 1924 (see below for synonymy)SK02 `--T. weymouthii (Müll.Hal.) Broth. 1924 [=Phascum weymouthii, Astomum weymouthii, Systegium weymouthii]SK02 RhamphidiumB57 |--R. montanum [=Didymodon montanus]OZB-O03 `--R. veitchiiB57 Nomina nuda: Triquetrella albicuspis Broth. ex Watts 1906SK02 Triquetrella triquetra Müll.Hal. ex Kindb. 1889SK02
Anoectangium aestivum (Hedw.) Mitt. 1869SK02 [=Gymnostomum aestivumOZB-O03, Pleurozygodon aestivusD24; incl. Anoectangium compactum f. brevifoliumOZB-O03, A. compactumS04, Gymnostomum compactumD24, Zygodon compactusOZB-O03]
Calyptopogon mnioides (Schwägr.) Broth. 1902 [=Streptopogon mnioides, Tortula mnioides; incl. Barbula crispata, Calyptopogon crispatus, Streptopogon crispatus, Tortula crispata, Barbula wilhelmii, Calyptopogon wilhelmii, Streptopogon wilhelmii, Tortula wilhelmii]SK02
Eucladium verticillatum (Brid.) Bruch & Schimp. in Bruch, Schimp. & Gümbel 1846S04 [=Weissia verticillata Hedw. ex Bridel 1801OZB-O03, Bryum verticillatum Linnaeus ex Withering 1801OZB-O03, Grimmia verticillataOZB-O03, Mollia verticillataD24]
Goniomitrium acuminatum ssp. enerve (Hooker & Wilson) Fife & Seppelt 2001 [=G. enerve Hooker & Wilson 1846, Physcomitrium enerve]SK02
Gymnostomum aeruginosum Smith 1804SK02 [=Mollia aeruginosaD24, Trichostomum aeruginosumOZB-O03; incl. Gymnostomum articulatumOZB-O03, G. rupestre f. compactum non G. compactumOZB-O03, G. rupestre f. gracilescensOZB-O03, Barbula incavataD24, G. rupestre f. ramosissimumOZB-O03, G. rupestreD24, Trichostomum rupestreOZB-O03, Weissia rupestrisS04, Gymnostomum rupestre f. stelligerumOZB-O03]
Gymnostomum calcareum Nees & Hornsch. in Nees et al. 1823S04 [=Hymenostylium calcareumOZB-O03, Mollia calcareaD24, Trichostomum calcareumOZB-O03; incl. Gymnostomum calcareum var. australeSK02, G. calcareum var. longifoliumSK02, G. rigidulumOZB-O03, Eucladium tasmanicumSK02]
Gymnostomum viridulum Brid. 1826S04 [=Weisia calcarea var. viridula non W. viridulaD24; incl. Gym. luisieriS04, Gyroweisia luisieriS04]
Gyroweisia tenuis (Schrad. ex Hedw.) Schimp. 1876S04 [=Gymnostomum tenueD24, Mollia tenuisD24, Weissia tenuisS04; incl. Gyroweisia tenuis var. acutifoliaOZB-O03, Gyr. tenuis var. badiaOZB-O03, Trichostomum reflexum var. gymnostomumOZB-O03]
Hennediella heimii (Hedw.) Zander 1993SK02 [=Desmatodon heimiiS04, Gymnostomum heimiiD24, Pottia heimii (Hedw.) Hampe 1837SK02; incl. G. affineOZB-O03, G. systyliumOZB-O03]
Hennediella stanfordensis (Steere) Blockeel 1990S04 [=Tortula stanfordensis Steere 1951SK02, Hyophila stanfordensisS04]
*Leptophascum leptophyllum (Müll. Hal.) Guerra & Cano 2000SK02, S04 [=Phascum leptophyllum Müll.Hal. 1888SK02, Chenia leptophylla (Müll.Hal.) Zander 1993SK02; incl. Tortula evanescens Broth. 1916SK02, T. rhizophylla (Sak.) Iwats. & Saito 1972SK02, T. vectensisS04]
Microbryum brevicaule (Taylor) Zander 1993 [=Gymnostomum brevicaule, Pottia brevicaulis (Taylor) Müll.Hal. 1849]SK02
Microbryum floerkeanum (Weber & Mohr) Schimp. 1860S04 [=Acaulon floerkeanumOZB-O03, Phascum floerkeanumS04; incl. Phascum floerkeanum var. badiumD24, OZB-O03]
Molendoa hornschuchiana (Hook.) Lindb. ex Limpr. 1886 [=Anoectangium hornschuchianum; incl. M. hornschuchiana var. tenuinervis]OZB-O03
Paraleptodontium recurvifolium (Taylor) Long 1982S04 [=Bryoerythrophyllum recurvifoliumS04, Bryum recurvifoliumD24, Didymodon recurvifoliusD24, Leptodontium recurvifoliumS04, Trichostomum recurvifoliumS04]
Pottiopsis caespitosa (Bruch ex Brid.) Blockeel & Sm. 1998S04 [=Pottia caespitosaS04, Trichostomum caespitosumS04, Weisia caespitosaD24]
Protobryum bryoides (Dicks.) Guerra & Cano 2000S04 [=*Mildeella bryoides (Dicks.) Limpricht 1885OZB-O03, Phascum bryoidesD24, Pottia bryoidesS04, Tortula protobryoides Zander 1993OZB-O03]
Splachnobryum obtusum (Brid.) Müll.Hal. 1869 [incl. S. baileyi Broth. 1888, S. geheebii Fleischer 1904, S. indicum Hampe & Müll.Hal. 1872]SK02
Tetrapterum brachypelma (Müll.Hal.) Broth. 1924 [=Phascum brachypelma, Astomum brachypelma, Systegium brachypelma]SK02
Tetrapterum cylindricum (Taylor) Jaeger 1869 [=Astomum cylindricum, Phascum tetragonum var. cylindricum, Systegium cylindricum, Tetrapterum capense var. cylindricum; incl. T. australe (nom. inv.)]SK02
Tetrapterum sullivanii (Müll.Hal.) Broth. 1924 [=Phascum sullivanii, Astomum sullivanii (preoc.), Pleuridium sullivanii, Systegium sullivanii]SK02
Tetrapterum tetrapteroides (Müll.Hal.) Broth. 1924 [=Phascum tetrapteroides, Astomum tetrapteroides, Systegium tetrapteroides]SK02
Trachycarpidium brisbanicum (Müll.Hal.) Stone 1975 [=Acaulon brisbanicum, Astomum brisbanicum, Sphaerangium brisbanicum; incl. As. wattsii]SK02
Triquetrella papillata (Hooker & Wilson) Broth. 1902 [=Didymodon papillatus, Leptodontium papillatum; incl. Triquetrella filiformis, Zygodon preissianus, T. preissiana (Hampe) Müll.Hal. 1897, Z. preissii (l. c.), Leskea rubricaulis, Triquetrella scabra, Zygodon scaber (n. n.)]SK02
Triquetrella tasmanica (Broth.) Granzow 1989 [=Anomodon tasmanicus Broth. 1900, Anoectangium tasmanicum (nom. inv.); incl. Triquetrella curvifolia]SK02
*Type species of generic name indicated
References
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[SK02] Streimann, H., & N. Klazenga. 2002. Catalogue of Australian Mosses. Flora of Australia Supplementary Series 17. Australian Biological Resources Study: Canberra.