Belongs within: Harpalinae.
Contains: Sphallomorpha, Adelotopus.
The Pseudomorphini are a group of often flattened ground beetles found living under bark. They are often found in association with ants and larvae of some species are physogastric inquilines in ant nests (Lawrence & Britton 1991). The majority of genera in this group are ovoviviparous (Baehr 2008). They may be related to the Graphipterini, a mostly African group with large eyes and long tibial spurs.
Characters (Lawrence & Britton 1991): Antennal scape not visible from above; antennae sometimes short and clavate; head with deep antennal sulcus between eye and mandible; legs more or less concealed beneath body, with may be flattened and laterally explanate.
<==Pseudomorphini [Pseudomorphinae]
| i. s.: Cryptocephalomorpha Ritsema 1874BB01
|--+--+--SphallomorphaOM08
| | `--Orthogonius [Orthogoniinae, Orthogoniini]OM08
| | |--O. brevilabrisB55
| | |--O. clarkeiB55
| | |--O. davidisB89
| | |--O. duplicatus [incl. O. alternans]B89
| | `--O. profundestriatusB89
| `--+--AdelotopusOM08
| |--Paussotropus Waterhouse 1877B08
| | `--P. cylindricus (Chaudoir 1862) (see below for synonymy)B08
| `--Cainogenion Notman 1925B08
| |--*C. ipsoides (Newman 1837) [=Adelotopus ipsoides]B08
| | |--C. i. ipsoidesB08
| | `--C. i. occidentale Baehr 1997B08
| |--C. creberrimum (Blackburn 1901) [=Adelotopus creberrimus]LM87
| |--C. ephippiatum (Newman 1856) [=Adelotopus ephippiatus]LM87
| `--C. obscurum (Castelnau 1867) [=Adelotopus obscurus; incl. A. subopacus Macleay 1871]LM87
`--+--Pseudomorpha Kirby 1825OM08, LM87 (see below for synonymy)
| | i. s.: P. angustataOM08
| | P. confusa Notman 1925LM87
| |--*P. (Pseudomorpha) excrucians Kirby 1825LM87
| |--P. (Austropseudomorpha Baehr 1997)BB01
| `--P. (Notopseudomorpha Baehr 1997)BB01
`--GraphipteriniBB01
|--Piezia Brullé 1834BB01
|--Trichopiezia Nègre 1955BB01
|--Discoptera Semenov 1889BB01
|--Corsyra Dejean 1825BB01
| `--C. fusulaC01
`--Graphipterus Latreille 1802OM08, L02
|--G. circumdatus Raffr. 1885G89
|--G. cordigerOM08
|--G. limbatusOM08
|--G. minutusB14
|--G. serrator (Forskål 1775)B14 [=Carabus serratorG89]
|--G. trilineatus [=Carabus trilineatus]L02
`--G. variegatus [=Carabus variegatus]L02
Paussotropus cylindricus (Chaudoir 1862) [=Adelotopus cylindricus, Cainogenion cylindricum; incl. *Paussotropus parellelus Waterhouse 1877]B08
Pseudomorpha Kirby 1825OM08, LM87 [incl. Axinophorus Dejean & Boisduval 1829BB01, Drepanus Dejan 1831BB01, Heteromorpha Kirby 1825 non Hübner 1822BB01]
*Type species of generic name indicated
References
[B08] Baehr, M. 2008. New and rare pseudomorphine species from Western Australia (Insecta, Coleoptera, Carabidae, Pseudomorphinae). Records of the Western Australian Museum 24 (2): 151–180.
[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.
[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.
[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 100–111.
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.
[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.
[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.
[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.