Psychodinae

Belongs within: Psychodoidea.
Contains: Telmatoscopus, Brunettia, Psychoda.

The Psychodinae, moth flies, are a group of densely hairy flies whose larvae are most commonly found in moist or semi-aquatic habitats (Wagner & Ibáñez-Bernal 2009).

Characters (from Wagner & Ibáñez-Bernal 2009): Eye reniform with extension towards midline forming eye bridge above antennal insertions; eyes continuous or separated, rarely with eye bridge short and acutely pointed; flagellomeres fusiform, pyriform or nodiform.

<==PsychodinaeWI-B09
    |--Maruina [Maruinini]WI-B09
    |    `--M. hirtaWM96
    |--PericominiWI-B09
    |    |--LepidiellaWI-B09
    |    `--Pericoma Walker 1856S53
    |         |--P. albipesS53
    |         |--P. (Clytocerus) americanaQV81
    |         |--P. bancrofti Tonnoir 1920S53
    |         |--P. confusa Satchell 1953S53
    |         |--P. pseudoalbipes Satchell 1953S53
    |         `--P. townsvillensis Taylor 1915S53
    |--ParamormiiniWI-B09
    |    |--Clogmia albipunctataWI-B09, WT11
    |    |--TelmatoscopusWI-B09
    |    `--Paramormia [incl. Duckhousiella]WI-B09
    |         |--P. furcataWI-B09
    |         `--P. ustulataWI-B09
    |--MormiiniWI-B09
    |    |--Atrichobrunettia Satchell 1953WI-B09, S53
    |    |    |--*A. alternata Satchell 1953S53
    |    |    `--A. subalternata Satchell 1953S53
    |    |--BryopharsosWI-B09
    |    |--Eugenys clavellataWI-B09
    |    |--MormiaO98
    |    `--BrunettiaWI-B09
    |--PsychodiniWI-B09
    |    |--Trichopsychoda Tonnoir 1922WI-B09, Q59
    |    |    |--*T. hirtella Tonnoir 1922Q59
    |    |    |--T. boninensis Quate 1959Q59
    |    |    |--T. carolinensis Quate 1959Q59
    |    |    |--T. insulicolaWI-B09, QV81
    |    |    `--T. montana Satchell 1953S53
    |    |--ThreticusWI-B09
    |    |--FeuerborniellaWI-B09
    |    |--PsychodaWI-B09
    |    `--PhilosepedonWI-B09
    |         |--P. helicisWM96
    |         |--P. labecula Quate 1963P92
    |         |--P. mauroae Wagner & Masteller 1996WM96
    |         |--P. oppositaWM96
    |         `--P. quateiWM96
    `--SetomiminiWI-B09
         |--Nemoneura Tonnoir 1929WI-B09, S53
         |    `--N. liparotesWI-B09
         |--Didicrum pyramidonWI-B09
         |--DesmiozaWI-B09
         |--Didimioza symphiliaWI-B09
         |--ThrysocanthusWI-B09
         |--ValeriannaWI-B09
         |--Neoarisemus maesiWI-B09
         |--NeurosystasisWI-B09
         |--Armillipora selvica Quate 1996WI-B09
         |--*Australopericoma caudata [incl. A. wirthi]WI-B09
         |--Balbagathis discuspisWI-B09
         |--SetomimaWI-B09
         |--PlatyplastinxWI-B09
         |--MicrommatosWI-B09
         |    |--M. simplexWI-B09
         |    `--M. sylvaticumWI-B09
         |--CaenobrunettiaWI-B09
         |    |--C. sarculosaWI-B09
         |    `--C. subditiciaWI-B09
         |--TonnoiraWI-B09
         |    |--T. cavernicolaWI-B09
         |    `--T. rapiformisWI-B09
         |--AlepiaWI-B09
         |    |--A. dioculaWI-B09
         |    |--A. hirtiventrisWI-B09
         |    `--A. longinoiWI-B09
         `--ArisemusWI-B09
              |--*A. boxiWM96
              |--A. amydrusWI-B09
              |--A. buzbyae Wagner & Masteller 1996WM96
              |--A. mariannae Wagner & Masteller 1996WM96
              `--A. waidei Wagner & Masteller 1996WM96

Psychodinae incertae sedis:
  Eurygarka helicisQV81
  BreviscapusQV81
  NeotelmatoscopusO98
  EpacretronO98
  GerobrunettiaO98
  Notiocharis Eaton 1912–1913O98, S53
    |--*N. insignisS53
    |--N. dimorpha Satchell 1953S53
    `--N. pallida Satchell 1953S53
  PeripsychodaO98
  UlomyiaRD77
  TermitomastusRD77
  TermitodipteronRD77
  TermitadelphosRD77
  Thornburghiella marginalisBW09 [=Pericoma marginalisQV81]

*Type species of generic name indicated

References

[BW09] Buck, M., N. E. Woodley, A. Borkent, D. M. Wood, T. Pape, J. R. Vockeroth, V. Michelsen & S. A. Marshall. 2009. Key to Diptera families—adults. In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 95–156. NRC Research Press: Ottawa.

[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[Q59] Quate, L. W. 1959. Diptera: Psychodidae. Insects of Micronesia 12 (4): 435–484.

[QV81] Quate, L. W., & J. R. Vockeroth. 1981. Psychodidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 293–300. Research Branch, Agriculture Canada.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10^th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[S53] Satchell, G. H. 1953. The Australian Psychodidae (Diptera), part I. Australian Journal of Zoology 1 (3): 357–418.

[WI-B09] Wagner, R., & S. Ibáñez-Bernal. 2009. Psychodidae (sand flies, and moth flies or owl flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 319–335. NRC Research Press: Ottawa.

[WM96] Wagner, R., & E. C. Masteller. 1996. New moth flies (Diptera: Psychodidae) and a key to species from Puerto Rico. Proceedings of the Entomological Society of Washington 98 (3): 450–464.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.