Ptomaphagini

Belongs within: Leiodidae.

The Ptomaphagini are a group of soil-dwelling scavenging beetles, some of which are found in subterranean habitats such as animal burrows or caves (Peck 2001).

Characters (from Peck 2001): Head with occipital carina, resting against front of pronotum when head is in repose; cervical sclerites present; elytra with glossy surface, setal bases arranged in strigae; procoxal cavities internally closed; posterior coxae contiguous; tarsi five-segmented; hind margin of hind tibia with two inner long spines and outer row or comb of short and equal spines only.

<==PtomaphaginiS05
    |--Adelopsis Portevin 1907P90
    |    |--A. bioforestae Salgado 2002S05
    |    |--A. brasiliensis Jeannel 1936S05
    |    |--A. coronaria Gnaspini & Peck 1996S05
    |    `--A. tandapi Salgado 2005S05
    `--Ptomaphagus Illiger 1798 (see below for synonymy)P01
         |  i. s.: P. consobrinusP90
         |         P. germariP92
         |         P. hirtus (Tellkampf 1844)B14
         |         P. shapardiP90
         |         P. tenuicornisLC20
         |         P. troglodytesLC20
         `--P. (Adelops Tellkampf 1884)S05, P01
              |--P. (A.) bordoni (Jeannel 1964) [=Adelopsis bordoni]S05
              `--P. (A.) sciakyi (Zoia 1992) [=Adelopsis sciakyi]S05

Ptomaphagus Illiger 1798 [incl. Appaladelopsis Gnaspini 1996, Echinocoleus Horn 1885, Merodiscus Jeannel 1934, Tupania Szymczakowski 1961]^P01

*Type species of generic name indicated

References

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[LC20] Lü, L., C.-Y. Cai, X. Zhang, A. F. Newton, M. K. Thayer & H.-Z. Zhou. 2020. Linking evolutionary mode to palaeoclimate change reveals rapid radiations of staphylinoid beetles in low-energy conditions. Current Zoology 66 (4): 435–444.

[P90] Peck, S. B. 1990. Insecta: Coleoptera Silphidae and the associated families Agyrtidae and Leiodidae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1113–1136. John Wiley & Sones: New York.

[P01] Peck, S. B. 2001. Leiodidae Fleming, 1821. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 250–258. CRC Press: Boca Raton.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[S05] Salgado, J. M. 2005. Cholevinae (Coleoptera, Leiodidae) from Ecuador: new data and two new species. Graellsia 61 (1): 51–60.