Pyxicephalidae

Belongs within: Diplasiocoela.

The Pyxicephalidae are an African clade of frogs united by molecular data (Frost et al. 2006).

<==Pyxicephalidae
    |  i. s.: Phrynopsis Pfeffer 1893 [Phrynopsinae]FG06
    |--Pyxicephalinae [Pyxicephalina]PW11
    |    |--Aubria Boulenger 1917FG06
    |    |    `--A. subsigillataPW11
    |    `--Pyxicephalus Tschudi 1838FG06
    |         |--P. adspersusPW11
    |         `--P. edulisPW11
    `--CacosterninaePW11
         |  i. s.: Nothophryne Poynton 1963FG06
         |         Ericabatrachus Largen 1991PW11, FG06
         |--Anhydrophryne Hewitt 1919PW11, FG06
         |    `--A. rattrayiPW11
         `--+--Tomopterna Duméril & Bibron 1841PW11, FG06 [Tomopterninae]
            |    |--T. natalensisPW11
            |    `--+--T. krugerensisPW11
            |       `--+--+--T. delalandiiPW11
            |          |  `--T. tuberculosaPW11
            |          `--+--+--T. lugangaPW11
            |             |  `--T. marmorataPW11
            |             `--+--T. damarensisPW11
            |                `--+--T. cryptotisPW11
            |                   `--T. tandyiPW11
            `--+--+--Strongylopus Tschudi 1838PW11, FG06
               |  |    |--*S. fasciatus (Burchell 1824) [=Rana fasciata]PR62
               |  |    `--S. bonaespeiPW11
               |  `--+--Poyntonia Channing & Boycott 1989PW11, FG06
               |     |    `--P. paludicolaPW11
               |     `--Cacosternum Boulenger 1887PW11, FG06
               |          |--+--C. nanumPW11
               |          |  `--+--C. boettgeriPW11
               |          |     `--C. capensePW11
               |          `--+--C. platysPW11
               |             `--Microbatrachella Hewitt 1926PW11, FG06
               |                  `--M. capensisPW11
               `--+--+--Natalobatrachus Hewitt & Methuen 1912PW11, FG06
                  |  |    `--N. bonebergiPW11
                  |  `--Arthroleptella Hewitt 1926FG06
                  |       |--+--A. lightfootiPW11
                  |       |  `--A. villiersiPW11
                  |       `--+--+--A. bicolorPW11
                  |          |  `--A. subvocePW11
                  |          `--+--A. drewesiiPW11
                  |             `--A. landdrosiaPW11
                  `--Amietia Dubois 1987PW11, FG06 [incl. Afrana Dubois 1992FG06]
                       |  i. s.: A. amieti (Laurent 1976) [=Rana (Afrana) amieti]FG06
                       |         A. desaegeri (Laurent 1972) [=Rana (Afrana) desaegeri]FG06
                       |         A. dracomontana (Channing 1978) [=Rana (Afrana) dracomontana]FG06
                       |         A. inyangae (Poynton 1966) [=Rana (Afrana) inyangae]FG06
                       |         A. johnstoni (Günther 1894) [=Rana (Afrana) johnstoni]FG06
                       |         A. lubricaFB17
                       |         A. ruwenzorica (Laurent 1972) [=Rana (Afrana) ruwenzorica]FG06
                       |         A. vandijki (Visser & Channing 1997) [=Rana (Afrana) vandijki]FG06
                       |--A. fuscigula (Duméril & Bibron 1841) [=Rana (Afrana) fuscigula]FG06
                       `--+--‘Strongylopus’ grayii (Smith 1849)PW11, PR62 [=Rana (Strongylopus) grayiiFG06]
                          `--+--A. angolensis (Bocage 1866) [=Rana (Afrana) angolensis]FG06
                             `--A. vertebralis [=Rana (Amietia) vertebralis]FG06

*Type species of generic name indicated

References

[FB17] Feng, Y.-J., D. C. Blackburn, D. Liang, D. M. Hillis, D. B. Wake, D. C. Cannatella & P. Zhang. 2017. Phylogenomics reveals rapid, simultaneous diversification of three major clades of Gondwanan frogs at the Cretaceous–Paleogene boundary. Proceedings of the National Academy of Sciences of the USA 114 (29): E5864–E5870.

[FG06] Frost, D. R., T. Grant, J. Faivovich, R. H. Bain, A. Haas, C. F. B. Haddad, R. O. de Sá, A. Channing, M. Wilkinson, S. C. Donnellan, C. J. Raxworthy, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. M. Green & W. C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1–370.

[PR62] Parker, H. W., & W. D. L. Ride. 1962. Rana fasciata Burchell, 1824 (Amphibia): proposed designation of a neotype under the plenary powers. Z.N.(S.) 1253. Bulletin of Zoological Nomenclature 19 (5): 290–292.

[PW11] Pyron, R. A., & J. J. Wiens. 2011. A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians. Molecular Phylogenetics and Evolution 61: 543–583.