Belongs within: Tephritoidea.
The Richardiidae are a group of flies, restricted to the New World, that mostly breed in decaying vegetable matter. They are divided between the subfamilies Epiplateinae, in which males have the phallus tip more or less simple and females have separate cerci on the ovipositor, and Richardiinae with an apically complex phallus and ovipositor cerci fused and adapted for piercing (Hancock 2010).
Characters (from Hancock 2010): Small to medium-sized (body length 3–15 mm). Head with single orbital seta. Stalk-eyes or genal extensions occasionally present. Frons may be considerably extended or swollen; occiput depressed above neck but occasionally enlarged behind eyes. Body colour yellow to brown or black, sometimes with darker or pruinose bands or stripes on scutum. Dorsum of thorax often setulose; thoracic chaetotaxy not extensive, presutural dorsocentrals absent. Scutellum with two or four marginal setae; if four then one pair basolateral. Postmesocoxal bridge often complete. Wings with one subcostal break, sometimes also with humeral break; cup closed apically by short sinuous or slightly convex vein. Wings frequently with apical spot, median band, basal and other markings. Legs often banded and usually bearing ventral femoral bristles on all or some legs. Two basal abdominal tergites fused, may be narrowed (pedunculate) or tapering basally more gradually. Usually strong preapical bristles present laterally on tergite 2. Occasionally with metallic blue or green abdomen contrasting with thorax. Male genitalia complex with long coiled phallus. Female with piercing-style ovipositor; two spermathecae present.
<==RichardiidaeH10 |--EpiplateinaeH10 | |--Schnusimyia parvulaH10 | |--EpiplateaH10 | | |--E. erosaS87 | | `--E. honduranaH10 | |--OmomyiaH10 | | |--O. hirsutaS87 | | `--O. regularisS87 | `--AutomolaH10 | |--A. atomariaH10 | |--A. calopteraH10 | `--A. rufaH10 `--RichardiinaeH10 |--Macrostenomyia gueriniiH10 |--Coniceps nigerH10 |--Oedematella czernyiH10 |--OzaeninaH10 |--OceaniciaH10 |--JohnrichardiaH10 |--AntineuromyiaH10 |--OdontomerellaH10 |--Hemixantha flavicornisH10 |--Setellida caerulescensH10 |--SpheneuolenaH10 |--Euolena egregiaH10 |--AcomphaH10 |--Neoidiotypa appendiculataH10 |--PanermyaH10 |--MegalothoracaH10 |--BatrachophthalmumH10 |--Poecilomyia longicornisH10 |--CladiscophlepsH10 |--BeebeomyiaH10 |--MaerorichardiaH10 |--RichardiodesH10 |--SepsisomaH10 | |--S. erythrocephalumH10 | `--S. flavescensH10 |--SetelliaH10 | |--S. apexH10 | |--S. costalisH10 | `--S. unispinosaH10 |--CoilometopiaH10 | |--C. costalisH10 | |--C. longicornisH10 | `--C. trimaculataH10 |--OdontomeraH10 | |--O. basistrigaH10 | |--O. ferrugineaH10 | |--O. fulvida [=Megamerina fulvida]MB10 | `--O. nitensH10 |--MelanolomaH10 | |--M. affinisH10 | |--M. canospilaH10 | |--M. decrepitaH10 | `--M. viatrixH10 `--RichardiaH10 |--R. eburneosignataH10 |--R. teevaniS87 | |--R. t. teevaniS87 | `--R. t. annulipesS87 `--R. telescopicaH10
*Type species of generic name indicated
References
[H10] Hancock, E. G. 2010. Richardiidae (richardiid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 871–879. NRC Research Press: Ottawa.
[MB10] Marshall, S. A., & M. Buck. 2010. Syringogastridae (syringogastrid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 843–846. NRC Research Press: Ottawa.
[S87] Steyskal, G. C. 1987. Richardiidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 833–837. Research Branch, Agriculture Canada.