Belongs within: Oestroidea.
Contains: Johnsoniini, Boettcheriini, Blaesoxipha, Sarcophagini.
The Sarcophaginae are a diverse lineage of flesh flies including necrophagous, coprophagous and parasitoid forms. Most have specific diets but the New World species Sarcodexia lambens has been bred from both live and dead animals and has been known to cause human myiasis. Oxysarcodexia, a Neotropical dung-breeding genus, is characterised by the presence in males ofan apical posteroventral ctenidium of flattened bristles on the mid femur (Pape & Dahlem 2010).
Characters (from Pape & Dahlem 2010): Antennal arista usually distinctly plumose; notopleuron usually with subprimary bristles; metasternal area usually setose; hind coxa usually setose posteriorly; coxopleural streak absent; male abdominal sternites 2–4 exposed and overlapping margins of corresponding tergites.
<==Sarcophaginae [Raviniini, Sarcodexiini]PD10 |--+--OxysarcodexiaKP10 [incl. Hybopygia Enderlein 1928L59] | | |--O. avuncula (Lopes 1933)KP10 | | |--O. cingarusS87 | | |--O. timidaPD10 | | `--O. varia (Walker 1836)MD13 (see below for synonymy) | `--+--Sinopiella rufopilosa Lopes & Tibana 1982KP10 | `--NotochaetiniS87 | |--Anolisimyia blakeaeS87 | `--Notochaeta Aldrich 1916KP10 | `--N. bufonivora [=Lepidodexia (Notochaeta) bufonivora]PD10 `--+--+--+--CucutomyiiniF92 | | `--JohnsoniiniKP10 | `--+--Tricharaea Thomson 1868KP10, L59 (see below for synonymy) | | |--T. brevicornis (Wiedemann 1830) (see below for synonymy)L59 | | `--T. occidua (Fabricius 1794)KP10 | `--SarothromyiiniS87 | |--Sarothromyia simplexS87 | |--Sarcophagula occiduaS87 | `--SarcofahrtiopsisKP10 | |--S. cuneata (Townsend 1935)KP10 | |--S. farriS87 | |--S. jamaicensisBW09 | |--S. paternaS87 | `--S. thryopteronthosPD10 `--+--+--HelicobiaKP10 | | | i. s.: H. morionellaPD10 | | |--H. rapax (Walker 1849)KP10 | | `--+--H. pilipleura Lopes 1939KP10 | | `--H. resinata (Hall 1933)KP10 | `--+--‘Tricharaea’ femoralis (Schiner 1868)KP10 | |--BoettcheriiniKP10 | `--EngelimyiaKP10 | |--E. bosqiPD10 | `--E. inops (Walker 1849)KP10 `--+--+--+--Dexosarcophaga transita Townsend 1917KP10 | | `--+--‘Blaesoxipha’ setosa (Salem 1938)KP10 | | `--ArgoraviniaKP10 [incl. RaviniopsisPD10] | | |--A. modestaS87 | | `--A. rufiventris (Wiedemann 1830)KP10 | `--Protodexiini [Blaesoxiphini, Servaisiini, Tephromyiini]S87 | | i. s.: Fletcherimyia fletcheriS87 | | MantidophagaS87 | | Opsophyto opiferaS87 | | Stenaulacotheca spatulataS87 | | Servaisia Robineau-Desvoidy 1863 (see below for synonymy)F92 | | |--S. erythrura (Meigen 1826) (see below for synonymy)F92 | | |--‘Acridiophaga’ aculeataS87 | | |--S. coloradensisS87 | | |--S. cothurnata (Hsue 1978) [=Blaesoxipha (Servaisia) cothurnata]F92 | | |--S. fossoria (Pandellé 1896)F92 | | |--S. jakovlevi Rohdendorf 1937F92 | | |--S. mixta (Rohdendorf 1928) [=Blaesoxipha mixta]F92 | | |--S. rossica (Villeneuve 1911) [=Blaesoxipha rossica]F92 | | `--S. subamericana (Rohdendorf 1932) [=Blaesoxipha (Servaisia) subamericana]F92 | | AmblycoryphenesS87 | | Agriella Villeneuve 1911 (see below for synonymy)F92 | | |--*A. pandellei Villeneuve 1911F92 | | `--A. gobica (Rohdendorf 1928) [=Blaesoxipha gobica]F92 | | Tephromyia Brauer & Bergenstamm 1891F92 | | Locustaevora Rohdendorf 1928F92 | |--BlaesoxiphaKP10 | `--+--Chlorosarcophaga Townsend 1919KP10 | `--SpirobolomyiaKP10 | |--S. flavipalpis (Aldrich 1916)KP10 | `--S. singularisS87 `--+--+--TitanogrypaKP10 | | |--T. larvicidaPD10 | | |--T. luculenta (Lopes 1938)KP10 | | `--T. placidaPD10 | `--Ravinia Robineau-Desvoidy 1863KP10 | |--*R. striata (Fabricius 1794) [=Musca striata; incl. Sarcophaga haematodes Meigen 1826]F92 | |--R. acerbaS87 | |--R. floridensisPD10 | `--R. querulaS87 `--+--SarcophaginiKP10 `--+--+--SarcodexiaKP10 | | |--S. innotaS87 | | `--S. lambens (Wiedemann 1830)KP10 | `--VillegasiaKP10 | |--V. almeidaiPD10 | `--V. postuncinata (Hall 1933)KP10 `--PeckiaKP10 |--+--P. intermutans (Walker 1861)KP10 | `--Peckiamyia minutipenis (Hall 1937)KP10 `--+--+--P. alvarengai Lopes & Tibana 1982KP10 | `--P. collusor (Curran & Walley 1934)KP10 `--+--P. chrysostoma (Wiedemann 1830)KP10 |--P. guloKP10 `--P. uncinata (Hall 1933)KP10 Sarcophaginae incertae sedis: TripanurgaPD10 NephochaetopteryxPD10 Panava inflataPD10 WulpiscaPD10 Pacatuba matthewsiPD10 TapacuraPD10 RetrocitomyiaPD10 Rettenmeyerina serrataPD10 Thomazomyia pilipesPD10 EmdenimyiaPD10 |--E. korytkowskiiPD10 `--E. meyersiPD10 MalacophagulaPD10 MalacophagomyiaPD10 ComasarcophaginiS87 |--Comasarcophaga texanaPD10 `--Mecynocorpus salvumS87 Udamopyga niagaranaPD10 Halliosca declinataPD10 Chrysagria duodecimpunctataPD10, S87 Oxyvinia panamensisPD10 HallimyiaPD10 Cistudinomyia cistudinis (Aldrich 1916)S87, K37 [=Sarcophaga cistudinisK37] Bellieria [Bellieriini]S87 `--B. melanuraS87 Cucullomyia pedunculataS87 TylomyiaS87 NeophytoiniS87 |--Neophyto setosaS87 `--Oppiopsis sheldoniS87 0--MicrocerellaPD10 [MicrocerelliniF92] | `--M. hypopygialisS87 `--+--SarcothromyiopsiniF92 `--+--EmblemasomaPD10 [incl. ColcondamyiaPD10, PessoamyiaPD10; EmblemasomatiniF92] | |--‘Colcondamyia’ auditrixS87 | `--E. erroPD10 `--ImpariiniF92 |--Speciosia speciosaS87 |--Abapa deletaS87 `--Kellymyia kellyiS87
Agriella Villeneuve 1911 [incl. Anablaesoxipha Villeneuve 1928, Asioblaeoxipha Rohdendorf 1937]F92
Oxysarcodexia varia (Walker 1836)MD13 [=Sarcophaga variaMD13, *Hybopygia variaL59; incl. S. milleri Johnston & Tiegs 1922MD13]
Servaisia Robineau-Desvoidy 1863 [incl. Acridiophaga Townsend 1917, Blaesoxiphomima Verves 1986]F92
Servaisia erythrura (Meigen 1826) [=Sarcophaga erythrura; incl. *Se. erythrocera Robineau-Desvoidy 1863]F92
Tricharaea Thomson 1868KP10, L59 [incl. Euleucomyia Townsend 1934L59, Leucoravinia Townsend 1928L59, Mallonotum Aldrich 1925L59, Xenagria Blanchard 1939L59]
Tricharaea brevicornis (Wiedemann 1830) [=Tachina brevicornis, *Mallonotum brevicornis; incl. Dexia albicans Walker 1854, Sarcophaga albicans, Tricharaea albicans, *Euleucomyia australis Townsend 1934, Sarcophaga canescens Thomson 1869, *Xenagria mallogaster Blanchard 1939, Liopyga pellita Townsend 1931, *Leucoravinia quadraticephala Townsend 1928, *Tricharaea scatophagina Thomson 1869]L59
*Type species of generic name indicated
References
[BW09] Buck, M., N. E. Woodley, A. Borkent, D. M. Wood, T. Pape, J. R. Vockeroth, V. Michelsen & S. A. Marshall. 2009. Key to Diptera families—adults. In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 95–156. NRC Research Press: Ottawa.
[F92] Fan Z. 1992. Key to the Common Flies of China 2nd ed. Science Press: Beijing.
[K37] Knipling, E. F. 1937. The biology of Sarcophaga cistudinis Aldrich (Diptera), a species of Sarcophagidae parasitic on turtles and tortoises. Proceedings of the Entomological Society of Washington 39 (5): 91–101.
[KP10] Kutty, S. N., T. Pape, B. M. Wiegmann & R. Meier. 2010. Molecular phylogeny of the Calyptratae (Diptera: Cyclorrhapha) with an emphasis on the superfamily Oestroidea and the position of Mystacinobiidae and McAlpine’s fly. Systematic Entomology 35: 614–635.
[L59] Lopes, H. de S. 1959. A revision of Australian Sarcophagidae (Diptera). Studia Ent. 2 (1–4): 33–67.
[MD13] Meiklejohn, K. A., M. Dowton, T. Pape & J. F. Wallman. 2013. A key to the Australian Sarcophagidae (Diptera) with special emphasis on Sarcophaga (sensu lato). Zootaxa 3680 (1): 148–189.
[PD10] Pape, T., & G. A. Dahlem. 2010. Sarcophagidae (flesh flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1313–1335. NRC Research Press: Ottawa.
[S87] Shewell, G. E. 1987. Sarcophagidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1159–1186. Research Branch, Agriculture Canada.