Satyrini

Belongs within: Satyrinae.
Contains: Heteronympha, Satyrus, Erebia, Euptychia.

The Satyrini are a diverse group of butterflies with larvae that feed mostly on grasses. Representatives include the heaths and ringlets of the Holarctic genus Coenonympha in which the antennal club is abrupt and the fore wings have vein 1A+2A and the cell greatly dilated towards the base. The Old World genus Ypthima has a narrow antennal club and only veins Sc and CuA are swollen towards the base in the fore wing (Braby 2000).

<==Satyrini
    |  i. s.: HeteronymphaWWN03
    |         CercyonisWWN03
    |           |--C. boopisS72
    |           |--C. oetusS72
    |           |--C. pegalaWWN03
    |           `--C. silvestrisS72
    |                |--C. s. silvestrisS72
    |                `--C. s. paulusS72
    |         Aphantopus hyperanthusWWN03
    |--+--DirinaJ03
    |  |--MelanargiaP01 [MelanargiinaJ03]
    |  |    `--M. japygia [incl. M. japygia var. suwarovia]P01
    |  `--+--PronophilinaJ03
    |     `--SatyrusJ03
    |--HypocystinaJ03
    |    |--ZipaetisJ03
    |    |--LamprolenisJ03
    |    |--XenicaJ03
    |    |    |--X. achantaO90
    |    |    |--X. correae Olliff 1890O90
    |    |    |--X. fulva Olliff 1890O90
    |    |    |--X. klugiiO90
    |    |    |--X. lathoniellaO90
    |    |    `--X. orichoraO90
    |    `--Hypocysta Westwood 1851B00
    |         |--H. adiante (Hübner 1831)B00
    |         |    |--H. a. adianteB12
    |         |    `--H. a. antirius Butler 1868B12
    |         |--H. angustata Waterhouse & Lyell 1914B00
    |         |--H. euphemia Westwood 1851B00
    |         |--H. irius (Fabricius 1775)B00
    |         |--H. metirius Butler 1875B00
    |         `--H. pseudirius Butler 1875B00
    `--+--ErebiaJ03
       |--+--ManiolaWWN03 [ManiolinaJ03]
       |  |    `--M. jurtinaWWN03
       |  `--CoenonymphaB66 [CoenonymphinaJ03]
       |       |--C. heroP01
       |       |--C. iphisP01
       |       |--C. oedipus [=Neonympha oedipus]B66
       |       |    |--C. o. oedipusI92
       |       |    `--C. o. annulifer Butler 1877I92
       |       |--C. pamphilusP01
       |       `--C. tulliaS72
       |            |--C. t. tulliaS72
       |            `--C. t. monoS72
       `--+--EuptychiaJ03
          `--YpthiminaJ03
               |--AustroypthimaJ03
               `--Ypthima Hübner 1818B00 [incl. Xois Hewitson 1865J03, B00]
                    |--Y. arctous (Fabricius 1775)B12 [=Xois arctousB00]
                    |--‘Xois’ fulvidaPP02
                    |--Y. granulosaB01
                    |--Y. masakii Ito 1947I92
                    |--Y. pusillaB00
                    |--Y. riukiuana Matsumura 1906I92
                    |--‘Xois’ sesaraPP02
                    `--Y. yayeyamana Nire 1920I92

*Type species of generic name indicated

References

[B00] Braby, M. F. 2000. Butterflies of Australia: their identification, biology and distribution vol. 2. CSIRO Publishing: Collingwood (Victoria).

[B12] Braby, M. F. 2012. The butterflies of El Questro Wilderness Park, with remarks on the taxonomy of the Kimberley fauna, Australia. Records of the Western Australian Museum 27 (2): 161–175.

[B66] Butler, A. G. 1866. A monograph of the genus Euptychia, a numerous race of butterflies belonging to the family Satyridae; with descriptions of sixty species new to science, and notes on their affinities &c. Proceedings of the Zoological Society of London 1866: 458–504.

[B01] Butler, A. G. 1901. An account of a collection of butterflies made by the Rev. K. St. Aubyn Rogers between Mombasa and the forests of Taveta. Annals and Magazine of Natural History, series 7, 7: 22–35.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[J03] Jong, R. de. 2003. Are there butterflies with Gondwanan ancestry in the Australian region? Invertebrate Systematics 17: 143–156.

[O90] Olliff, A. S. 1890. On Rhopalocera from Mt. Kosciusko, New South Wales. Proceedings of the Linnean Society of New South Wales, series 2, 4 (3): 619–624.

[PP02] Patrick, B., & H. Patrick. 2002. Butterflies of Fiji. Weta 24: 5–12.

[P01] Pável, J. 1901. Lepkék [Lepidopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 171–177. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[S72] Shields, O. 1972. Flower visitation records for butterflies (Lepidoptera). Pan-Pacific Entomologist 48 (3): 189–203.

[WWN03] Wahlberg, N., E. Weingartner & S. Nylin. 2003. Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea). Molecular Phylogenetics and Evolution 28: 473–484.