Belongs within: Conjunctae.
Contains: Laccopterum, Scaraphites, Conopterum, Neocarenum, Monocentrum, Euryscaphus, Carenum, Scarites, Clivina, Dyschiriodes, Dyschirius.
The Scaritinae are a diverse group of ground beetles with body forms commonly modified for digging. The body is pedunculate with a marked constriction between pro- and mesothorax. The fore coxal cavities are closed behind and the mid coxal cavities laterally open, partially closed by the mesepimeron (Lawrence & Britton 1991). The tribe Clivinini includes relatively small forms with a prominent, setiform unguitractor plate on the tarsus whereas members of the Scaritini have a much smaller plate and tend to be much larger in overall size (Ball & Bousquet 2001).
<==Scaritinae [Scarititae, Scrobifera]B14 |--Migadopini [Migadopinae]R-JC01 | |--Migadops latusR-JC01 | `--Stichonotus Sloane 1910LB91, LM87 | |--*S. leai Sloane 1910LM87 | |--S. limbatus Sloane 1915LM87 | `--S. piceus Sloane 1915LM87 |--Scaritini [Scaritides]B1990 | |--EuryscaphusLB91 | |--CarenumLB91 | |--ScaritesBB01 | `--Pasimachus Bonelli 1813BB01 | | i. s.: P. elongatus LeConte 1846BB01 | | P. subangulatus (Chaudoir 1862)B14 | `--P. (Emydopterus Lacordaire 1854)BB01 `--Clivinini [Clivinides]BB01 |--ClivinaBB01 |--DyschiriodesBB01 |--DyschiriusS96 |--Oxydrepanus Putzeys 1866BB01 | `--O. rufus (Putzeys 1846) [incl. O. brevicarinatus Putzeys 1846]FS90 |--Halocoryza Alluaud 1919BB01 | `--H. arenaria (Darlington 1939)BB01 |--Clivinarchus Sloane 1896S96 | `--*C. perlongus Sloane 1896LM87 |--Steganomma Macleay 1887S96, LM87 | `--*S. porcatum Macleay 1887LM87 |--Ardistomis Putzeys 1846BB01 | |--A. atripennis Putz. 1867FS90 | `--A. laevistriata Fleutiaux & Sallé 1890FS90 |--Aspidoglossa Putzeys 1846BB01 | |--A. semicrenata Chd. 1843 [incl. A. guadeloupensis Putz. 1846]FS90 | `--A. subangulata (Chaudoir 1843)BB01 |--Semiardistomis Kult 1950BB01 | |--S. puncticollis (Dejean 1831)BB01 | `--S. viridis (Say 1823)BB01 |--Akephorus LeConte 1852BB01 | |--A. marinus LeConte 1852BB01 | `--A. obesus LeConte 1866B14 `--Schizogenius Putzeys 1846BB01 | i. s.: S. freyi Baehr 1983BB01 `--S. (Genioschizus Whitehead 1972)BB01 `--S. (G.) crenulatus LeConte 1852BB01 Scaritinae incertae sedis: Ochyropus gigasB55 TaeniolobusB55 |--T. laevipennisB55 |--T. lucidusB55 | |--T. l. lucidusB55 | `--T. l. strigicepsB55 `--T. vilhenai Basilewsky 1955B55 Trilophus congoanusB55 Syleter Andrewes 1941 [=Psilus Putzeys 1877 nec Panzer 1806 nec Fischer de Waldheim 1813]LM87 |--*S. paradoxus (Putzeys 1868) [=Ardistomis paradoxa]LM87 |--S. andrewesi (Basilewsky 1931) [=Dyschirius andrewesi, Psilus andrewesi]B55 |--S. congoensisB55 |--S. papua Darlington 1962LM87 `--‘Clivina’ trapezicollis Bates 1889 [=Psilus trapezicollis]B1890 Pseudoclivina Kult 1947B55 `--P. grandisB55 Rhysocara Sloane 1916LM87 `--*R. crassa Sloane 1916LM87 Platysphyrus Sloane 1905LM87 `--*P. tibialis Sloane 1905LM87 LaccopterumLM87 Philoscaphus Macleay 1871LM87 |--*P. tuberculatus (Macleay 1863) [=Carenum tuberculatum]LM87 |--P. barnardi Macleay 1888LM87 |--P. bicostatus Sloane 1905LM87 |--P. carinatus (Macleay 1864) [=Carenum carinatum]LM87 |--P. costalis Macleay 1873 (see below for synonymy)LM87 `--P. mastersii Macleay 1871 [incl. P. duboulayi Blackburn 1890]LM87 ScaraphitesLM87 ConopterumLM87 NeocarenumLM87 Trichocarenum Blackburn 1892LM87 |--*T. elderi Blackburn 1892LM87 |--T. castelnaui Sloane 1905LM87 `--T. cylindricum Sloane 1897LM87 Neoscaphus Sloane 1888LM87 `--*N. simplex Sloane 1888LM87 Epilectus Blackburn 1888 [=Carenarchus Sloane 1905, Eurygnathus Blackburn 1888 non Wollaston 1854]LM87 |--*E. fortis (Blackburn 1888) [=*Eurygnathus fortis, *Carenarchus fortis]LM87 `--E. mastersi (Macleay 1869) [=Neocarenum mastersi]LM87 MonocentrumLM87 Geoscaptus Chaudoir 1855LM87 |--*G. laevissimus Chaudoir 1855 (see below for synonymy)LM87 |--G. cacus (Macleay 1863) (see below for synonymy)LM87 |--G. crassus Sloane 1895LM87 |--G. macleayi Chaudoir 1879LM87 `--G. plicatulus (Castelnau 1867) [=Scarites plicatulus]LM87 OmoglymmiusB14 |--O. americanus (Laporte 1836)B14 `--O. hamatusB14 Solenogenys funkei Adis 1981B14 Distichus planus (Bonelli 1813)B89, G89 [=Scarites planusG89]
Geoscaptus cacus (Macleay 1863) [=Scarites cacus; incl. S. approximatus Macleay 1865, S. bipunctatus Castelnau 1867, S. mitchellii Castelnau 1867, S. ruficornis Castelnau 1867, S. subporcatulus Macleay 1865, S. substriatus Castelnau 1867, S. waterhousei Macleay 1865]LM87
*Geoscaptus laevissimus Chaudoir 1855 [incl. Scarites bostocki Castelnau 1867, S. damastes Macleay 1863, S. geryon Macleay 1863, S. jacksoniensis Macleay 1865, S. planiusculus Macleay 1865]LM87
Philoscaphus costalis Macleay 1873 [incl. P. crassus Blackburn 1888, P. lateralis Macleay 1873, P. tepperi Blackburn 1888]LM87
*Type species of generic name indicated
References
[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.
[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.
[B89] Bates, H.-W. 1889. Contributions a la faune Indo-Chinoise. 3e mémoire. Carabidae. Annales de la Société Entomologique de France, 6e série 9: 261–286.
[B1890] Bates, H. W. 1890. Rectification suivante. Annales de la Société Entomologique de France, 6e série 9: ccxl.
[B1990] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.
[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.
[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).
[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.
[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.
[S96] Sloane, T. G. 1896. On the Australian Clivinides (fam. Carabidae). (Revision of the Australian species of the genus Clivina with the description of a new genus, Clivinarchus). Proceedings of the Linnean Society of New South Wales 21 (2): 143–257.