Belongs within: Bibionomorpha.
The Scatopsoidea are a clade of flies united by wings with M1+2 fused to Rs for a distance, and M3 and im lost; at least living species also have the larval head greatly reduced (Blagoderov et al. 2002). Members of the Scatopsidae have a single-segmented palpus and five to ten antennal flagellomeres (Amorim 2009) whereas Synneuridae have a four-segmented palpus and ten to fourteen flagellomeres (Peterson & Cook 1981). The Protoscatopsidae are early members of the Scatopsoidea known from the Jurassic period (Blagoderov et al. 2002).
<==ScatopsoideaBLM02 |--MesoscatopseRJ93 [ProtoscatopsidaeBLM02] | `--M. rohdendorfi Kovalev in Kalugina & Kovalev 1985RJ93 |--Synneuridae [Canthyloscelidae, Hyperoscelididae]BLM02 | |--Exiliscelis californiensisPC81 | |--Prohyperoscelis jurassicus Kovalev in Kalugina & Kovalev 1985RJ93 | `--SynneuronPC81 | |--S. annulipesPC81 | |--S. decipiensPC81 | `--S. silvestrePC81 `--ScatopsidaeA09 | i. s.: Procolobostema Cook 1971P92, H02 | |--P. hurdi Cook 1971P92 | |--P. incisa Cook 1971P92 | |--P. longicorne Cook 1971P92 | `--P. obscurum Cook 1971P92 | Aldrorardiella Enderlein 1936H02 |--Ectaetia [Ectaetiinae]A09 | `--E. clavipesC81 |--AspistinaeA09 | |--Arthria analisA09, C81 | `--AspistesA09 | |--A. berolinensisG20 | `--A. hartiA09 |--PsectrosciarinaeA09 | |--Anapausis solutaA09 | `--PsectrosciaraA09 | |--P. californicaA09 | `--P. serrataA09 `--ScatopsinaeA09 | i. s.: Sinoscatopse Hong 2002H02 | `--*S. eocenica Hong 2002H02 |--ColobostematiniA09 | |--Colobostema variatumA09 | |--BorneoscatopseA09 | |--Holoplagia guamensisA09 | |--LumpuriaA09 | |--Cookella albitarsisA09 | `--FerneiellaA09 |--ScatopsiniA09 | |--Reichertella Enderlein 1912A09, H02 | |--Apiloscatopse uncinataA09 | |--PharsoreichertellaA09 | `--Scatopse Greoffroy 1762H02 | |--S. bilaminata Cook 1971P92 | |--S. fuscipesCM91 | |--S. latrinarumR26 | |--S. nigraL02 | |--S. notata [=Bibio (Scatopse) notatus]G20 | `--S. primula Cook 1971P92 |--RhegmoclematiniA09 | |--Diamphidicus [Diamphidicina]A09 | |--Holoclema [Holoclematina]A09 | |--RhegmoclematinaA09 | | |--ParmaferiaA09 | | |--Thripomorpha truncatum [=Rhegmoclema truncatum]A09 | | `--Rhegmoclema Enderlein 1936H02 | `--ParascatopsinaA09 | |--Parascatopse wirthiA09 | |--AustrocleminaA09 | |--Neorhegmoclemina Cook 1955A09, H02 | | `--N. asymmetricaA09 | `--Rhegmoclemina Enderlein 1936A09, H02 | `--R. bimaculataA09 `--SwammerdamelliniA09 |--Akorhexoza cactivoraA09 |--Coboldia fuscipesA09 |--PararhexosaA09 |--HawomersleyaA09 |--AbrhexozaA09 |--CookaA09 |--BrahemyiaA09 |--Swammerdamella Enderlein 1912H02 | |--S. obtusaA09 | `--S. prima Cook 1971P92 |--QuateiellaA09 | |--Q. inexpectataA09 | `--Q. minusculaA09 `--RhexozaA09 |--R. incisaA09 `--R. similisA09
*Type species of generic name indicated
References
[A09] Amorim, D. de S. 2009. Scatopsidae (minute black scavenger flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 347–355. NRC Research Press: Ottawa.
[BLM02] Blagoderov, V. A., E. D. Lukashevich & M. B. Mostovski. 2002. Order Diptera Linné, 1758. The true flies (=Muscida Laicharting, 1781). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 227–240. Kluwer Academic Publishers: Dordrecht.
[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).
[C81] Cook, E. F. 1981. Scatopsidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 313–319. Research Branch, Agriculture Canada.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[H02] Hong Y. 2002. Amber Insect of China. Beijing Scientific and Technological Publishing House.
[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.
[PC81] Peterson, B. V., & E. F. Cook. 1981. Synneuridae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 321–324. Research Branch, Agriculture Canada.
[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.
[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.
[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.