Belongs within: Schizophora.
Contains: Conopidae, Sciomyzidae.
The Sciomyzoidea are a group of acalyptrate flies characterised by having the face desclerotised along the vertical midline, and possessing a densely setose frontal vitta. Members include the Coelopidae, robust flies with a somewhat flattened habitus most commonly found along coastlines (Vockeroth 1987). The Dryomyzidae are a Holarctic group with a large and prominent clypeus bulging below the lower margin of the face in profile. The coastal Helcomyzidae resembly Dryomyzidae but have a triangular prosternum forming a precoxal bridge. The Helosciomyzidae, found in Australasia and South America, resemble members of the Sciomyzidae but for an anteriorly protruding prelabrum, a short second antennal segment, spaced anterior spines on the costa, and two strong sternopleural bristles present (Colless & McAlpine 1991). Stylogaster is a genus of slender flies, females of which have an elongate ovipositor usually about as long as the rest of the abdomen, that develop as endoparasites of orthopteroids such as cockroaches and crickets (Skevington et al. 2010).
<==Sciomyzoidea [Helcomyzinae]WT11 |--+--Natalimyza [Natalimyzidae]WT11 | `--Heterocheila [Heterocheilidae]WT11 | |--H. buccataWT11 [=Oedoparea buccataS87] | `--H. hannaiS87 `--+--Stylogaster [Stylogastrinae]WT11 | |--S. biannulataSP87 | |--S. malgachensisA71 | |--S. neglectaSTC10 | `--S. stylosusSTC10 `--+--+--+--Pelidnoptera [Phaeomyiidae, Phaeomyiinae]WT11 | | | `--P. nigripennisWT11 | | `--+--ConopidaeWT11 | | `--SciomyzidaeWT11 | `--Dryomyzidae [Dryomyzinae]WT11 | |--Oedoparena glaucaWT11 | |--ParactoraS87 | |--MaorimyiaS87 | |--Palaeotimia lhoestiS87 | |--Prodryomyza electricaS87 | `--DryomyzaWT11 | |--D. anilisWT11 | `--D. flaveolaS87 `--+--+--Huttonina [Huttoninidae, Huttonininae]WT11 | | `--H. glabraWT11 | `--HelcomyzaWT11 [incl. ActoraRD77; Helcomyzidae] | |--H. mirabilisWT11 | `--H. ustulataS87 `--+--Helosciomyzidae [Helosciomyzinae]WT11 | |--Neosciomyza luteipennisWT11 | |--HelosciomyzaCM91 | |--EurotocusCM91 | |--XenosciomyzaCM91 | |--PolytocusCM91 | `--SciogriphoneuraS87 `--Coelopidae [Phycodromidae]WT11 | i. s.: Dasycoelopa australisCM91 | Chaetocoelopa sydneyensisCM91 | MalacomyiaV87 | Baeopterus robustusV87, N91 | Icaridion debileV87, KP10 |--Lopa [Lopinae]M10 | `--L. convexaWT11 `--CoelopinaeM10 |--GluminiM10 | |--Gluma nitidaKP10 | `--Coelopina anomalaM10 `--Coelopa [Coelopini]M10 | i. s.: C. alluaudi [incl. C. palauensis]O98 | C. littoralisO81 | C. stejnegeriV87 |--C. (Fucomyia)M10 | |--C. (F.) frigidaM10 | `--C. (F.) nebularumM10 `--C. (Neocoelopa) vanduzeeiM10
*Type species of generic name indicated
References
[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.
[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).
[KP10] Kutty, S. N., T. Pape, B. M. Wiegmann & R. Meier. 2010. Molecular phylogeny of the Calyptratae (Diptera: Cyclorrhapha) with an emphasis on the superfamily Oestroidea and the position of Mystacinobiidae and McAlpine’s fly. Systematic Entomology 35: 614–635.
[M10] Mathis, W. N. 2010. Coelopidae (kelp flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1013–1016. NRC Research Press: Ottawa.
[N91] Norris, K. R. 1991. General biology. In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 68–108. Melbourne University Press: Carlton (Victoria).
[O81] O’Brien, C. O. 1981. A. A. Book of New Zealand Wildlife: A guide to the native and introduced animals of New Zealand. Lansdowne Press: Auckland.
[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.
[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.
[STC10] Skevington, J. H., F. C. Thompson & S. Camras. 2010. Conopidae (thick-headed flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 847–855. NRC Research Press: Ottawa.
[SP87] Smith, K. G. V., & B. V. Peterson. 1987. Conopidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 749–756. Research Branch, Agriculture Canada.
[S87] Steyskal, G. C. 1987. Dryomyzidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 923–926. Research Branch, Agriculture Canada.
[V87] Vockeroth, J. R. 1987. Coelopidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 919–922. Research Branch, Agriculture Canada.
[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.