Sciophilini

Belongs within: Mycetophilidae.

The Sciophilini is a group of sciophiline fungus gnats distinguished by the presence of macrotrichia on the wings (Tonnoir & Edwards 1927). However, monophyly of the group has been questioned on the basis of similarities of some genera to species lacking macrotrichia (Vockeroth 1981).

Characters (from Tonnoir & Edwards 1927): Wing with macrotrichia on the membrane; microtrichia sometimes absent. Postnotum usually with hairs or bristles; Sc nearly always long; last section of R₁ several times as long as r-m, which is oblique; 7th abdominal segment usually large and visible.

<==Sciophilini
    |--TrizygiaTE27
    |--Loicia basifurcaV81
    |--AzanaV81
    |--Monoclona rufilateraV81
    |--CluzobraV81
    |--AcnemiaV81
    |--Polylepta guttiventrisV81
    |--AnaclileiaV81
    |--Baeopterogyna nudipesV81
    |--Neuratelia sayiV81
    |--Leptomorphus nebulosusV81
    |--Eudicrana obumbrataV81
    |--Neotrizygia Tonnoir & Edwards 1927TE27
    |    `--*N. obscura Tonnoir in Tonnoir & Edwards 1927TE27
    |--Morganiella Tonnoir & Edwards 1927TE27
    |    `--*M. fusca Tonnoir in Tonnoir & Edwards 1927TE27
    |--PhthiniaV81
    |    |--P. longiventris Tonnoir in Tonnoir & Edwards 1927TE27
    |    `--P. tanypusV81
    |--Taxicnemis Tonnoir & Edwards 1927TE27
    |    |--*T. ‘hirta’ (Marshall 1896) (see below for synonymy)TE27
    |    `--T. flava Edwards in Tonnoir & Edwards 1927TE27
    |--ParatiniaV81
    |    |--P. montana Plassmann 1976P76a
    |    `--P. recurvaV81
    |--SyntemnaV81
    |    |--S. hungaricaP76b
    |    |--S. nitidulaP76b
    |    `--S. vernalisV81
    |--AllocotoceraV81 [incl. Euryceras Marshall 1896TE27]
    |    |--A. anaclinoides (Marshall 1896) [=*Euryceras anaclinoides]TE27
    |    |--A. cephasi Edwards in Tonnoir & Edwards 1927TE27
    |    |--A. crassipalpis Tonnoir in Tonnoir & Edwards 1927TE27
    |    |--A. dilatata Tonnoir in Tonnoir & Edwards 1927TE27
    |    `--A. pulchellaV81
    |--Sciophila [incl. Megalopelma]V09
    |    |--S. adamsiP76b
    |    |--‘Megalopelma’ glabanumV81
    |    |--S. hirta Meigen 1818TC04
    |    |--S. novataV81
    |    |--S. ocreata Philippi 1865TC04
    |    |--S. parviareolata Santos Abreu 1920TC04
    |    `--S. striata [=Mycetophila (Sciophila) striata]G20
    |--AphelomeraTE27
    |    |--A. elongata Tonnoir in Tonnoir & Edwards 1927TE27
    |    |--A. forcipata Edwards in Tonnoir & Edwards 1927TE27
    |    |--A. longicauda Edwards in Tonnoir & Edwards 1927TE27
    |    |--A. majuscula Edwards in Tonnoir & Edwards 1927TE27
    |    |--A. marshalli Edwards in Tonnoir & Edwards 1927TE27
    |    |--A. opaca Tonnoir in Tonnoir & Edwards 1927TE27
    |    `--A. skusei Marshall 1896TE27
    |--Parvicellula Marshall 1896TE27
    |    |--*P. triangula Marshall 1896TE27
    |    |--P. apicalis Tonnoir in Tonnoir & Edwards 1927TE27
    |    |--P. fascipennis Edwards in Tonnoir & Edwards 1927TE27
    |    |--P. gracilis Tonnoir in Tonnoir & Edwards 1927TE27
    |    |--P. hamata Edwards in Tonnoir & Edwards 1927TE27
    |    |--P. nigricoxa Tonnoir in Tonnoir & Edwards 1927TE27
    |    |--P. obscura Tonnoir in Tonnoir & Edwards 1927TE27
    |    |--P. ruficoxa Tonnoir in Tonnoir & Edwards 1927TE27
    |    `--P. subhamata Tonnoir in Tonnoir & Edwards 1927TE27
    |--StenophragmaSS51
    `--Aneura Marshall 1896TE27
         |--*A. boletinoides Marshall 1896TE27
         |--A. appendiculata Tonnoir in Tonnoir & Edwards 1927TE27
         |--A. bispinosa Edwards in Tonnoir & Edwards 1927TE27
         |--A. defecta Edwards in Tonnoir & Edwards 1927TE27
         |--A. fagi (Marshall 1896) [=Sciophila fagi]TE27
         |--A. filiformis Tonnoir in Tonnoir & Edwards 1927TE27
         |--A. fusca Tonnoir in Tonnoir & Edwards 1927TE27
         |--A. longicauda Tonnoir in Tonnoir & Edwards 1927TE27
         |--A. longipalpis Tonnoir in Tonnoir & Edwards 1927TE27
         |--A. nitida Tonnoir in Tonnoir & Edwards 1927TE27
         `--A. pallida Edwards in Tonnoir & Edwards 1927TE27

*Taxicnemis ‘hirta’ (Marshall 1896) [=Sciophila hirta non Meigen 1818; incl. T. hirta var. bivittata Tonnoir & Edwards 1927]^TE27

*Type species of generic name indicated

References

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[P76a] Plassmann, E. 1976a. Zwei neue Pilzmücken aus dem Kaukasus (Diptera: Mycetophilidae). Senckenbergiana Biologica 57 (1–3): 69–70.

[P76b] Plassmann, E. 1976b. Vierter Nachtrag zu der Mycetophiliden-Sammlung des Senckenberg-Museums Frankfurt am Main (Diptera; Mycetophilidae). Senckenbergiana Biologica 57 (1–3): 77–79.

[SS51] Shaw, F. R., & M. M. Shaw. 1951. Relationships of certain genera of fungus gnats of the family Mycetophilidae. Smithsonian Miscellaneous Collections 117 (3): 1–23.

[TC04] Toft, R. J., & P. J. Chandler. 2004. Three introduced species of Mycetophilidae (Diptera: Sciaroidea) established in New Zealand. New Zealand Entomologist 27: 43–49.

[TE27] Tonnoir, A. L., & F. W. Edwards. 1927. New Zealand fungus gnats (Diptera, Mycetophilidae). Transactions and Proceedings of the New Zealand Institute 57: 747–878.

[V81] Vockeroth, J. R. 1981. Mycetophilidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 223–246. Research Branch, Agriculture Canada.

[V09] Vockeroth, J. R. 2009. Mycetophilidae (fungus gnats). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 267–278. NRC Research Press: Ottawa.