Scolytus

Belongs within: Scolytinae.

Scolytus is a genus of phloeophagous beetles found in Eurasia, northern Africa and the Americas. The greater number of species are monogamous with a smaller number of bigynous species, mostly found in the Neotropics (Wood 1986).

Characters (from Wood 1986): Eye oval, entire; antenna with funicle 7-segmented, club rather large, flattened; pronotum unarmed, lateral margins costate; scutellum depressed, subtriangular, apically (posteriorly) pointed; elytral bases depressed in scutellar area, appearing emarginate in median area; costal margin of elytra normal (straight) and overlapping metepisternum; protibia unarmed on lateral margin, outer apical angle extending into spine curving toward and exceeding inner apical angle, socketed denticles never present; metapleural suture descending subvertically to groove receiving groove on costal margin of elytra then turning abruptly and continuing parallel to groove to near metacoxal process; abdomen flexed upward from anterior margin of segment 2.

<==Scolytus Geoffroy 1762W86 (see below for synonymy)
    |--*S. scolytusW86 (see below for synonymy)
    |--S. claviger [=*Archaeoscolytus claviger]W86
    |--S. confusus [=Eccoptogaster confusus, *Confusoscolytus confusus]W86
    |--S. intricatus [=Eccoptogaster intricatus, *Tubuloscolytus intricatus]W86
    |--S. laevisK68
    |--S. maliS00
    |--S. marawitzi [=*Pinetoscolytus marawitzi]W86
    |--S. multistriatus (Marsham 1802)B14 (see below for synonymy)
    |--S. praecepsM38
    |--S. pygmaeus [=Bostrichus pygmaeus, *Pygmaeoscolytus pygmaeus]W86
    |--S. rugulosus [=Bostrichus rugulosus, *Ruguloscolytus rugulosus]W86
    |--S. schevryrewiB14
    `--S. ventralisWL09

Scolytus Geoffroy 1762^W86 [=Coptogaster Illiger 1807^W86, Eccoptogaster Gyllenhal 1813^W86, Ekkoptogaster Herbst 1793^W86; incl. Archaeoscolytus Butovitsch 1929^W86, Confusoscolytus Tsai & Huang 1962^A02, Pinetoscolytus Butovitsch 1929^W86, Pygmaeoscolytus Butovitsch 1929^W86, Ruguloscolytus Butovitsch 1929^W86, Scolytochelus Reitter 1913^W86, Spinuloscolytus Butovitsch 1929^W86, Tubuloscolytus Butovitsch 1929^W86]

Scolytus multistriatus (Marsham 1802)^B14 [=Ips multistriatus^W86, Eccoptogaster multistriatus^R13, *Scolytochelus multistriatus^W86, *Spinuloscolytus multistriatus^W86]

*Scolytus scolytus^W86 [=Bostrichus scolytus^W86, *Coptogaster scolytus^W86, *Eccoptogaster scolytus^W86, *Ekkoptogaster scolytus^W86, Hylesinus scolytus^L02; incl. Eccoptogaster destructor^R13, Hylesinus (Scolytus) destructor^{G20, S00}]

*Type species of generic name indicated

References

[A02] Anderson, R. S. 2002. Curculionidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 722–815. CRC Press.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[K68] Kinn, D. N. 1968. A new species of Pleuronectocelaeno (Acarina: Celaenopsidae) associated with bark beetles in North and Central America. Acarologia 9 (2): 191–205.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[M38] Muesebeck, C. F. W. 1938. The genus Dendrosoter Wesmael in the United States (Hymenoptera: Braconidae). Proceedings of the Entomological Society of Washington 40 (9): 281–287.

[R13] Reuter, O. M. 1913. Lebensgewohnheiten und Instinkte der Insekten bis zum Erwachen der sozialen Instinkte. R. Friedländer & Sohn: Berlin.

[S00] Siddiqi, M. R. 2000. Tylenchida: Parasites of plants and insects 2^nd ed. CABI Publishing: Wallingford (UK).

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

[W86] Wood, S. L. 1986. A reclassification of the genera of Scolytidae (Coleoptera). Great Basin Naturalist Memoirs 10: 1–126.