
Belongs within: Pentatomoidea.
The Scutelleridae, jewel bugs, are a group of mostly shiny and/or brightly coloured shield bugs with a very large scutellum mostly covering the abdomen dorsally, and a hamus (spur-vein) present in the basal cell of the hind wing (Carver et al. 1991). Members include the cotton harlequin bug Tectocoris diophthalmus, a significant pest of cotton bolls.
The Elvisurinae are large brown or grey species with strongly sulcate thoracic sterna. The Scutellerinae are brightly coloured with the hind margin of the pronotum truncate or slightly concave (Gross 1975).
<==Scutelleridae
|--Tetyra Fabricius 1803 [incl. Macraulax Dallas 1851; Tetyrinae]G75
| `--T. bipunctatus (Herrich Schaeffer 1839) [=Pachycoris bipunctatus, *Macraulax bipunctatus]G75
|--Tectocoris [Tectocorinae]CGW91
| |--T. cyanipes Fabr. 1803 (see below for synonymy)D14
| `--T. diophthalmus Kirk 1908C91, D14
|--OdontotarsinaeCGW91
| |--OdontotarsusL49
| | |--O. caudatus [=Pachycoris caudatus, Bellocoris caudatus; incl. Odontotarsus productus]L49
| | `--O. grammicus (see below for synonymy)L49
| |--EurygasterG75
| | |--E. hottentotus [=Tetyra hottentota, Scutellera hottentota]L49
| | |--E. integricepsRD77
| | |--E. maura (Linnaeus 1758)H01, L58 (see below for synonymy)
| | `--E. nigrocucullatusG75
| `--Morbora Distant 1899G75
| |--*M. australis Distant 1899G75
| |--M. hirtula Bergroth 1904G75
| `--M. schoutedeni Bergroth 1904G75
|--ElvisurinaeCGW91
| |--ElvisuraG75
| |--Solenostethium lynceumG75, L49 [=Tetyra lynceaL49, Coeloglossa lynceuL49; incl. Scutellera bilunata Lefebv. 1827L49]
| |--Solenotichus Martin 1897G75
| | `--S. circuliferus (Walker 1867) [=Sphaerocoris circuliferus; incl. *So. brevipes]G75
| |--Austrotichus Gross 1975G75
| | `--*A. rugosus Gross 1975G75
| `--Coleotichus White 1839G75
| | i. s.: C. nigrovarius Walker & Schouteden 1905K08
| | C. sordidus Walker & Schouteden 1904K08
| |--C. (Coleotichus)G75
| | |--*C. (C.) costatus (Fabricius 1787) [=Cimex costatus]G75
| | `--C. (C.) blackburniG75
| `--C. (Paracoleotichus) discrepansG75
`--ScutellerinaeCGW91
|--Scutiphora pedicellataCGW91, C91
|--Lampromicra senatorCGW91, C91
|--Calliphara Germar 1839G75
| |--*C. nobilis (Linnaeus 1763)G75 [=Cimex nobilisG75, Pentatoma nobilisL02]
| |--C. imperialisCGW91
| `--C. praslinia (Guérin 1830) [=Scutellera praslinia]G75
|--Choerocoris Dallas 1851G75
| |--*C. paganus (Fabricius 1775) [=Cimex paganus]G75
| `--C. variegatusG75
| |--C. v. variegatusG75
| `--C. v. similis Distant 1899G75
`--ScutelleraB35
|--S. cyanipoda Boisduval 1835B35
|--S. globusR26
|--S. lineataG20
|--S. pagana [=Tetyra pagana]B35
`--S. tongae Boisduval 1835B35
Scutelleridae incertae sedis:
PhiliaC70
|--P. elegans (Montrouz. 1861) [=Callidea elegans]D14
`--P. senatorC70
Chrysocoris sexmaculatus (Leach 1815) [=Scutellera sexmaculata]D14
PeocilocorisB14
Cantao parentumWEE70
Odontoscelis fuliginosaRD77, H01 (see below for synonymy)
Homaemus aeneifronsE66
RaphigasterL49
|--R. punctipennisL49 (see below for synonymy)
`--R. purpuripennis [=Pentatoma purpuripennis]L49
TrigonosomaL49
|--T. desfontainii [=Tetyra desfontainii]L49
`--T. nigellae [=Tetyra nigellae]L49
PsacastaL49
|--P. pedemontana [=Tetyra pedemontana, Ventocoris pedemontana]L49
`--P. tuberculata [=Tetyra tuberculata]L49
Podops inunctus [=Tetyra inuncta, Cimex inunctus]L49
Arctocoris maculiventris (Germ. 1839) [=Pachycoris maculiventris, Irochrotus maculiventris; incl. P. hirta]L49
Eurygaster maura (Linnaeus 1758)H01, L58 [=Cimex (Scutellatus) maurusL49, L58, Bellocoris maurusL49, Odontotarsus maurusL49, Pentatoma mauraL02, Scutellera mauraL49; incl. Cimex pictaL49, Eurygaster maura var. pictaH01]
Odontoscelis fuliginosaRD77, H01 [=Cimex fuliginosusL49, Arctocoris fuliginosusL49, Scutellera fuliginosaL49, Tetyra fuliginosaL49, Ursocoris fuliginosusL49; incl. T. dorsalisL49, U. dorsalisL49, T. lyturaL49, U. lyturusL49]
Odontotarsus grammicus [=Cimex grammicus, Pachycoris grammicus, Tetyra grammica; incl. Cimex purpureolineatus, Bellocoris purpureolineatus]L49
Raphigaster punctipennisL49 [=Cimex punctipennisL49; incl. Cimex (Rotundatus) griseus Linnaeus 1758L49, L58, Pentatoma griseaL02, Raphigaster griseusL49]
Tectocoris cyanipes Fabr. 1803 [=T. lineola var. cyanipes; incl. T. lineola var. banksi Don 1805, Cimex lineola Fabr. 1781 non Linnaeus 1758, T. lineola, T. lineola var. obliquus Walk. 1867, T. lineola var. tongae Boisd. 1832]D14
*Type species of generic name indicated
References
[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d’Urville. Faune entomologique de l’océan Pacifique, avec l’illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).
[C70] CSIRO. 1970. The Insects of Australia: A textbook for students and research workers. Melbourne University Press.
[C91] CSIRO. 1991. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1. Melbourne University Press: Carlton (Victoria).
[D14] Distant, W. L. 1914. Rhynchota from New Caledonia and the surrounding islands. In: Sarasin, F., & J. Roux (eds) Nova Caledonia: Forschungen in Neu-Caledonian und auf den Loyalty-Inseln. A. Zoologie vol. 1 pt 4 pp. 369–390, pls 11–12. C. W. Kreidels Verlag: Wiesbaden.
[E66] Evans, H. E. 1966. The Comparative Ethology and Evolution of the Sand Wasps. Harvard University Press: Cambridge (Massachusetts).
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[G75] Gross, G. F. 1975. Plant-feeding and Other Bugs (Hemiptera) of South Australia. Heteroptera—Part I. Handbook of the Flora and Fauna of South Australia.
[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.
[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.
[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis revised 10th ed. vol. 1. Laurentii Salvii: Holmiae.
[L49] Lucas, H. 1849. Exploration Scientifique de l’Algérie pendant les années 1840, 1841, 1842 publiée par ordre du gouvernement et avec le concours d’une commission académique. Sciences physiques. Zoologie. II. Histoire naturelle des animaux articulés. Troisième partie. Insectes (suite). Imprimerie Nationale: Paris.
[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.
[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.
[WEE70] Woodward, T. E., J. W. Evans & V. F. Eastop. 1970. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 387–457. Melbourne University Press.