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Belongs within: Cyphophthalmi.
The Sironidae are a Holarctic family of mite-like harvestmen with circular spiracles, simple claws, and single cheliceral dentition.
<==Sironidae [Sironides]
|--Suzukielus Juberthie 1970GS10, BG04
| `--*S. sauteri (Roewer 1916) [=Miopsalis sauteri]G00
`--+--+--Tranteeva Kratochvíl 1958BG04 [=Trenteeva (l. c.)G00]
| | `--*T. paradoxa Kratochvíl 1958 [=Trenteeva (l. c.) paradoxa]G00
| `--+--Siro Latreille 1797GS10, K18 [incl. Geosiro Rafalski 1958G00]
| | |--*S. rubens Latreille 1804G00, KMS24
| | |--S. balticus Dunlop & Mitov 2011PP19
| | |--S. beschkovi Mitov 1994BKG05
| | |--S. carpaticus Rafalski 1956GS10 [=S. (Geosiro) carpaticusG00]
| | |--S. clousi Giribet & Shear 2010GS10
| | |--S. coccineusR26
| | |--S. crassus Novak & Giribet 2006GS10
| | |--S. ligiae Giribet in Giribet et al. 2017KM20
| | |--S. platypedibus Dunlop & Giribet 2003PP19
| | |--S. richarti Benavides & Giribet in Giribet et al. 2017KM20
| | `--S. valleorum Chemini 1990GS10
| `--+--+--Holosiro Ewing 1923GS10
| | | `--*H. acaroides Ewing 1923 [=Siro acaroides]GS10
| | |--‘Siro’ calaveras Giribet & Shear 2010GS10
| | `--‘Siro’ sonoma Shear 1980BG04
| `--+--‘Siro’ shasta Giribet & Shear 2010GS10
| `--+--Neosiro Newell 1943GS10
| | |--*N. kamiakensis Newell 1943 [=Siro kamiakensis]GS10
| | `--N. exilis (Hoffman 1963)KMS24
| `--+--‘Siro’ boyerae Giribet & Shear 2010GS10
| `--‘Siro’ exilis Hoffman 1963GS10
`--+--+--Odontosiro Juberthie 1961BG04
| | `--*O. lusitanicus Juberthie 1962G00, KMS24
| `--ParasironidaeKMS24
| |--Parasiro Hanson & Sørensen 1904GS10, BG04
| | `--*P. corsicus (Simon 1872)G00, BG04 [=Cyphophthalmus corsicusG00, Siro corsicusG00]
| |--Cimmerosiro Karaman, Mitov & Snegovaya 2024KMS24
| | |--*C. krivolutskyi Karaman, Mitov & Snegovaya 2024KMS24
| | |--C. juberthiei Karaman, Mitov & Snegovaya 2024KMS24
| | `--C. rhodiensis Karaman, Mitov & Snegovaya 2024KMS24
| |--Tirrenosiro Karaman, Mitov & Snegovaya 2024KMS24
| | |--*T. minor (Juberthie 1958) [=Parasiro minor]KMS24
| | `--T. axeli Karaman, Mitov & Snegovaya 2024KMS24
| `--Ebrosiro Karaman, Mitov & Snegovaya 2024KMS24
| `--E. coiffaiti (Juberthie 1956) [=Parasiro coiffaiti]KMS24
`--+--+--Marwe Shear 1985BG04, G00
| | `--*M. coarctata Shear 1985G00
| `--+--Iberosiro de Bivort & Giribet 2004BG04
| | |--*I. distylos de Bivort & Giribet 2004BG04
| | `--I. rosae Giribet, Merino-Sáinz & Benavides in Giribet et al. 2017KM20
| `--Paramiopsalis Juberthie 1962GS10, G00
| |--*P. ramulosus Juberthie 1962G00
| |--P. anadonae Giribet, Merino-Sáinz & Benavides in Giribet et al. 2017KM20
| `--P. ramblae Benavides & Giribet in Giribet et al. 2017KM20
`--Cyphophthalmus Joseph 1868GS10, KMS24
| i. s.: C. ere Karaman 2008K08 [=Siro ere Karaman et al. 1994 (n. n.)G00]
| C. montenegrinus (Hadži 1973)BKG05 [=Siro montenegrinusG00]
| C. noctiphilus (Kratochvíl 1940)BKG05 [=Siro noctiphilusG00]
| C. ohridanus (Hadži 1973)BKG05 [=Siro ohridanusG00]
| C. silhavyi (Kratochvíl 1938)BKG05 [=Siro silhavyiG00]
|--+--C. eratoae (Juberthie 1968)BC07, BKG05 [=Siro eratoaeG00]
| `--C. gjorgjevici (Hadži 1933)BKG05 (see below for synonymy)
| |--C. g. gjorgjeviciBKG05
| `--C. g. klisurae (Hadži 1973)BKG05 [=Siro gjorgjevici klisuraeG00]
`--+--C. serbicus (Hadži 1973)BKG05 [=Siro serbicusG00]
`--+--*C. duricorius Joseph 1868G00, BKG05 [=Siro duricoriusBG04; incl. S. cyphopselaphus Joseph 1882G00]
| |--C. d. duricoriusBKG05
| |--C. d. bithynicus (Gruber 1969)BKG05 [=Siro duricorius bithynicusG00]
| |--C. d. bolei (Hadži 1973)BKG05 [=Siro duricorius boleiG00]
| |--C. d. corfuanus (Kratochvíl 1938)BKG05 [=Siro duricorius corfuanusG00]
| `--C. d. yalovensis (Gruber 1969)BKG05 [=Siro duricorius yalovensisG00]
`--+--C. minutus (Kratochvíl 1938)BKG05 [=Siro minutusG00]
`--C. teyrovskyi (Kratochvíl 1938)BKG05 [=Siro teyrovskyiG00]
Nomina nuda: Cyphophthalmus gordani Boyer, Karaman & Giribet 2005BKG05
Cyphophthalmus markoi Boyer, Karaman & Giribet 2005BKG05
Cyphophthalmus ognjanovici Boyer, Karaman & Giribet 2005BKG05
Cyphophthalmus rumijae Boyer, Karaman & Giribet 2005BKG05
Cyphophthalmus trebinjanum Boyer, Karaman & Giribet 2005BKG05
Siro martensi Karaman et al. 1994G00 [=Cyphophthalmus martensiBKG05]
Siro zetae Karaman et al. 1994G00 [=Cyphophthalmus zetaeBKG05]
Cyphophthalmus gjorgjevici (Hadži 1933)BKG05 [=Siro gjorgjeviciG00, S. dordevici (l. c.)G00, S. gjordjevici (l. c.)G00, S. gjorjevici (l. c.)G00]
*Type species of generic name indicated
References
[BG04] Bivort, B. L. de, & G. Giribet. 2004. A new genus of cyphophthalmid from the Iberian Peninsula with a phylogenetic analysis of the Sironidae (Arachnida: Opiliones: Cyphophthalmi) and a SEM database of external morphology. Invertebrate Systematics 18: 7–52.
[BC07] Boyer, S. L., R. M. Clouse, L. R. Benavides, P. Sharma, P. J. Schwendinger, I. Karunarathna & G. Giribet. 2007. Biogeography of the world: a case study from cyphophthalmid Opiliones, a globally distributed group of arachnids. Journal of Biogeography 34 (12): 2070–2085.
[BKG05] Boyer, S. L., I. Karaman & G. Giribet. 2005. The genus Cyphophthalmus (Arachnida, Opiliones, Cyphophthalmi) in Europe: a phylogenetic approach to Balkan Peninsula biogeography. Molecular Phylogenetics and Evolution 36: 554–567.
[G00] Giribet, G. 2000. Catalogue of the Cyphophthalmi of the world (Arachnida, Opiliones). Revista Ibérica de Aracnología 2: 49–76.
[GS10] Giribet, G., & W. A. Shear. 2010. The genus Siro Latreille, 1796 (Opiliones, Cyphophthalmi, Sironidae), in North America with a phylogenetic analysis based on molecular data and the description of four new species. Bulletin of the Museum of Comparative Zoology 160 (1): 1–33.
[K08] Karaman, I. M. 2008. A new Odiellus species from Serbia (Opiliones, Phalangiidae). In: S. E. Makarov, & R. N. Dimitrijević (eds) Advances in Arachnology and Developmental Biology. Papers Dedicated to Prof. Dr. Božidar P. M. Ćurčić pp. 275–279. Inst. Zool., Belgrade.
[KMS24] Karaman, I. M., P. G. Mitov & N. Snegovaya. 2024. Parasironidae fam. nov., a Cimmerian lineage of Mediterranean Cyphophthalmi (Opiliones), with the description of three new genera and four new species. European Journal of Taxonomy 921: 173–209.
[K18] Kury, A. B. 2018. Familial nomina in harvestmen (Arachnida, Opiliones). Bionomina 13: 1–27.
[KM20] Kury, A. B., A. C. Mendes, L. Cardoso, M. S. Kury & A. A. Granado. 2020. WCO-Lite: Online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0—Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018. Published by the author: Rio de Janeiro.
[PP19] Palencia, L., E. Peñalver, C. E. Prieto & F. J. Poyato-Ariza. 2019. First fossil harvestmen (Arachnida: Opiliones) from Spain and notes on the fossil record of Opiliones. Palaeontologica Electronica 22.1.5A: 1–18.
[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.