Belongs within: Sciomyzidae.
The Tetanocerini are a group of snail-killing flies whose larvae are active predators of terrestrial or aquatic mollusks. Adults are characterised by a head with an antero-dorsally projecting profile, a pedicel subequal to or twice the length of the first flagellomere, and absence of a proepisternal seta (Marinoni & Knutson 2010).
<==Tetanocerini [Tetanocerinae]MK10
|--Limnia unguicornisK87, K01
|--SepedoninusMK10
|--Sepedonella nanaMK10
|--Euthycera arcuataMK10
|--TetanoceroidesMK10
|--Anticheta testaceaK87
|--Poecilographa decoraK87
|--Dictyacium ambiguumK87
|--Pherbecta limenitisK87
|--Trypetoptera canadensisK87
|--Hedria mixtaK87
|--ThecomyiaMK10
| |--T. chrysacraMK10
| `--T. longicornisMK10
|--SepedomerusMK10
| |--S. bipuncticepsMK10
| `--S. macropusMK10
|--ElgivaK87
| |--E. rufaK01
| `--E. solicitaK87
|--HoplodictyaMK10
| |--H. setosaK87
| `--H. spinicornisMK10
|--TeutoniomyiaMK10
| |--T. costaricensisMK10
| `--T. plaumanniMK10
|--GuatemaliaMK10
| |--G. nigritarsisMK10
| `--G. straminataMK10
|--SepedoneaMK10
| |--S. guatemalanaMK10
| |--S. isthmiMK10
| `--S. lagoaMK10
|--ProtodictyaMK10
| |--P. brasiliensisMK10
| |--P. iguassuMK10
| `--P. nubilipennisMK10
|--DictyaMK10
| |--D. bergiMK10
| |--D. expansaK87
| |--D. guatemalanaMK10
| |--D. matthewsiK87
| `--D. umbrarumMK10
|--TetanoceraWT11
| |--T. algira Macquart 1847E12
| |--T. ferrugineaWT11
| |--T. griseicollisK87
| |--T. marginata [=Oscinis (Tetanocera) marginata]G20
| |--T. montanaK87
| |--T. phyllophoraK87
| |--T. planifronsR26
| |--T. plebejaK87
| |--T. plumosaMK10
| |--T. punctulataK01
| |--T. reticulataK01
| |--T. spretaMK10
| |--T. unipunctata Macquart 1847E12
| |--T. validaK87
| |--T. vicinaE66
| `--T. vittigeraK01
`--Sepedon [Sepedoninae]MK10
| i. s.: S. knutsoniMK10
| S. palustrisR26
| S. relictus van der Wulp 1897FO72
| S. rufipes [=Oscinis (Sepedon) rufipes]G20
| S. tenuipesS68
|--S. caeruleaFO72
|--S. fuscipennisFO72
|--S. macropusFO72
|--S. pacifica Cresson 1914FO72
|--S. praemiosa Giglio-Tos 1893FO72
|--S. spinipesFO72
| |--S. s. spinipesK87
| `--S. s. americanaK87
|--S. tenuicornisFO72
|--S. trinidadensisFO72
|--S. armipes groupFO72
| |--S. anchistaFO72
| |--S. armipesS68
| |--S. bifidaFO72
| |--S. capelleiFO72
| |--S. haplobasis Steyskal 1960FO72
| |--S. melanderiFO72
| `--S. pseudarmipesFO72
|--S. lindneri groupFO72
| |--S. guatemalanaFO72
| `--S. lindneriFO72
`--S. pusilla groupFO72
|--S. borealisFO72
|--S. lignatorFO72
|--S. neiliFO72
`--S. pusillaFO72
*Type species of generic name indicated
References
[E66] Evans, H. E. 1966. The Comparative Ethology and Evolution of the Sand Wasps. Harvard University Press: Cambridge (Massachusetts).
[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.
[FO72] Fisher, T. W., & R. E. Orth. 1972. Resurrection of Sepedon pacifica Cresson and redescription of Sepedon praemiosa Giglio-Tos with biological notes (Diptera: Sciomyzidae). Pan-Pacific Entomologist 48 (1): 8–20.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[K01] Kertész, K. 1901. Legyek [Dipteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 179–201. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[K87] Knutson, L. V. 1987. Sciomyzidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 927–940. Research Branch, Agriculture Canada.
[MK10] Marinoni, L., & L. Knutson. 2010. Sciomyzidae (snail-killing flies, marsh flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1017–1024. NRC Research Press: Ottawa.
[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.
[S68] Steyskal, G. C. 1968. The number and kind of characters needed for significant numerical taxonomy. Systematic Zoology 17 (4): 474–477.
[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.