Belongs within: Cerithiimorpha.
Contains: Melanoides.
The Thiaridae are a cosmopolitan family of freshwater caenogastropods. In the past, pretty much all lake-dwelling cerithioids have been treated as Thiaridae (or as Melaniidae, an older but invalid name for the same family) but this larger Thiaridae is now regarded as polyphyletic. Similarly, many unrelated freshwater gastropods with high-spired shells have historically been included in the genus Melania, an objective junior synonym of the genus Thiara. Thiaridae sensu stricto have a fringed mantle border; a small radula with a plain central tooth; and a paucispiral operculum with baso-columellar nucleus (Pilsbry & Bequaert 1927).
<==Thiaridae [Melaniidae]
|--Stomatopsis Stache 1871BR05 [Stomatopsidae, StomatopsinaeBR17]
| `--*S. cosinensis Stache 1871BR17
`--ThiarinaeBR17
|--MelanoidesBR17
`--Thiara Röding 1798BR05 (see below for synonymy)
| i. s.: ‘Melania’ abbreviata Defrance 1823F27
| ‘Melania’ abendanoniB50
| ‘Melania’ bulbosaC64
| ‘Melania’ cancellataC42
| ‘Melania’ corporosa [=M. (Striatella) corporosa]PB27
| ‘Melania’ decollata Reeve 1859PB27
| ‘Melania’ ebeninaPB27
| ‘Melania’ exiguaC64
| ‘Melania’ fasciolataF66
| ‘Melania’ ferrea Reeve 1859PB27
| ‘Melania’ gemmiferaB50
| ‘Melania’ hongkongensisPB27
| ‘Melania’ insulaesacraeB50
| ‘Melania’ lalemaeB50
| ‘Melania’ lateritiaF66
| ‘Melania’ mahalonensisB50
| ‘Melania’ mahalonicaB50
| ‘Melania’ masapensisB50
| ‘Melania’ maurula Reeve 1859PB27
| ‘Melania’ menkeanaC64
| ‘Melania’ neglecta Anthony 1854BC01
| ‘Melania’ newberryiC64
| ‘Melania’ nigrina Lea 1856C64
| ‘Melania’ occataC64
| ‘Melania’ palicolarumB50
| ‘Melania’ pallidula Reeve 1860 [=M. pallens]PB27
| ‘Melania’ patriarchalisB50
| ‘Melania’ phlebotomum Reeve 1859PB27
| ‘Melania’ plicifera [incl. M. silicula]C64
| ‘Melania’ sarasinorumB50
| ‘Melania’ scipio [incl. M. buschiana]C64
| ‘Melania’ semicancellataPB27
| ‘Melania’ shastaensis Lea 1856 [incl. M. shortaensis]C64
| ‘Melania’ tominangensisB50
| ‘Melania’ towutensisB50
| ‘Melania’ towuticaB50
| ‘Melania’ wahlamatensisC64
| ‘Melania’ warderianaC64
|--*T. (Thiara) amarula (Linnaeus 1758)BR17 (see below for synonymy)
| |--T. a. amarulaPB27
| `--‘Melania’ a. coacta [incl. M. crenularis v. Martens 1860, Tiara vouamica Bourguignat 1889]PB27
`--T. (Plotiopsis)BRW98
|--‘Plotiopsis’ australisPH10
|--T. (P.) balonnensisBRW98
`--T. (P.) lirata (Benson 1836)BRW98, TC89 (see below for synonymy)
Thiaridae incertae sedis:
Goniobasis Lea 1862W77
|--G. athleta [incl. G. glauca]B68a
|--G. bulbosa Gould 1847 [incl. G. newberryi Lea 1860]R63
|--G. curreyanaB68a
| |--G. c. curreyanaB68a
| `--G. c. lyoni Lea 1863 [incl. G. viridicata Lea 1863]B68a
|--G. depygisW79
|--G. floridensisH64
|--G. interioris Goodrich 1944R63
|--G. laqueataB68a
|--G. laurae Goodrich 1944R63
|--G. livescensW79
|--G. milesiiW79
|--G. nebrascensis Meek & Hayden 1856W77
|--G. nigrina (Lea 1856) (see below for synonymy)R63
|--G. proximaD84
|--G. pulchellaW79
|--G. semicarinata [incl. G. indianensis Pilsbry 1903]B68b
|--G. tenera (Hall 1845) [=Cerithium tenerum]W77
|--G. tenuicarinata (Meek & Hayden 1857) [=Melania tenuicarinata]W77
`--G. virginicaW79
Carbonispira Yen 1949TTE93, KC60
`--*C. scotica Yen 1949KC60
Horea Bourguignat 1888PB27
`--‘Melania’ tanganyicensis Smith 1880 [incl. *Horea tanganikana Bourguignat 1888]PB27
*Antimelania variabilis [=Melania variabilis]PB27
Juramelanoides Bandel 1991B91
`--*J. villersense (de Loriol 1865) [=Cerithium villersense]B91
Lavansia Bandel 1991B91
`--*L. mojonia Bandel 1991B91
RipalaniaBRW98
SermylasmaBRW98
Sermyla retractaPH10
Goniobasis nigrina (Lea 1856) [incl. G. californica (Clessin 1882), G. circumlineata Tryon 1865, G. draytonii Lea 1862]R63
Thiara Röding 1798BR05 [=Amarula Sowerby 1842PB27, Hydrognoma Gistel 1848PB27, Lithoparches Gistel 1848PB27, Melacantha Swainson 1840PB27, Melania Lamarck 1799BR05, Melanidia Rafinesque 1815PB27, Spirilla Humph. in Gray 1847PB27, Tiara Hermannsen 1849 non Swainson 1831PB27; incl. Melas Denys de Montfort 1810PB27; Melaniacea, Melanianae, Thiareae, Thiarinae, Tiarinae] BR05
Thiara (Plotiopsis) lirata (Benson 1836)BRW98, TC89 [=Melania (Tarebia) lirataTC89; incl. M. flavidaTC89, M. semigranosaTC89]
*Thiara (Thiara) amarula (Linnaeus 1758)BR17 [=Helix amarulaBR17, *Amarula amarulaPB27, *Hydrognoma amarulaPB27, *Lithoparches amarulaPB27, *Melacantha amarulaPB27, *Melania amarulaBR17, *Melanidia amarulaPB27, *Spirilla amarulaPB27, *Tiara amarulaPB27; incl. *Melas melanus Denys de Montfort 1810PB27]
*Type species of generic name indicated
References
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[PB27] Pilsbry, H. A., & J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of Congo malacology. Bulletin of the American Museum of Natural History 53 (2): 69–602, pls 10–77.
[PH10] Pinder, A. M., S. A. Halse, R. J. Shiel & J. M. McRae. 2010. An arid zone awash with diversity: patterns in the distribution of aquatic invertebrates in the Pilbara region of Western Australia. Records of the Western Australian Museum Supplement 78 (1): 205–246.
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