Belongs within: Cleroidea.
Contains: Temnochilini.
The Trogossitidae are a morphologically diverse group of predatory or mycophagous beetles, commonly found under bark. Members of the subfamily Egoliinae have a compact, one- or two-segmented antennal club whereas that of Trogossitinae is large, loose and three-segmented (Lawrence & Britton 1991). Members of the genus Calitys have a symmetrical antennal club and the dorsal surfaces of elytra are ornamented with costae and tubercles (Leschen 2002). The Decamerinae have toothed claws and feed on pollen (Leschen 2002). The Protopeltinae are small mycophagous species endemic to New Zealand.
Characters (from Lawrence & Britton 1991): Mostly ovate to elongate and more or less flattened, with prognathous head, short antennae bearing large, loose, slightly asymmetrical, 3-segmented club (1- to 2-segmented and compact in Egoliinae), and vestiture of short or long hairs, bristles or scales. Tarsal formula 5-5-5, first segment may be reduced and partly concealed by tibial apex. Abdomen with five ventrites, none connate. Larvae elongate, parallel sided, slightly to strongly flattened, lightly sclerotised, except for head and usually additional sclerites on thorax and T9, mandible usually with 1 or several hyaline processes at base of mesal edge; ventral mouth-parts consolidated, without articulating areas; hypostomal rods usually long and parallel; para-gular sclerites usually present between gula and hypostomal rods; urogomphi usually well developed.
<==Trogossitidae [Nemosomidae, Temnochilidae, Trogositidae]
|--DecamerinaeL02
|--ProtopeltinaeL02
|--EgoliinaeLB91
| |--Egolia variegataLB91
| `--Necrobiopsis tasmanicusB74
|--Calitys Thomson 1859 [incl. Nosodes LeConte 1861, Peltidea Motschulsky 1858 non Ach. 1803 (ICBN); Calitinae]L02
| |--C. minorB14
| `--C. scabra (Thunberg 1784)B14
`--TrogossitinaeB14
| i. s.: PhanodestaB14
| KolibaciaB14
| LepidopterixLB91
| |--L. decorataLB91
| |--L. nigrosparsaWH02
| `--L. turbataLB91
| Corticotomus Sharp 1891 [incl. Parafilumis Casey 1916]L02
| Cylidrella Sharp 1891L02
| `--C. championi Wickham 1916L02
| Euschaefferia Leng 1920 [incl. Pseudocotomus Van Dyke 1944, Stenodema Schaeffer 1918]L02
|--TemnochiliniL89a
|--NemosominiL89a
| |--Nemosoma Latreille 1804L02 [incl. HolocephalaL89e, Pseudalindria Fall 1910L02]
| | `--N. elongatumLC40 [=Cis (Nemosoma) elongatumG20; incl. Colydium fasciatumLC40]
| `--NomosomiaL89a
| |--N. simoni Lév. 1889L89a
| `--N. voraxL89a
`--Leperina [Gymnochilini]B14
| i. s.: L. conspicua Oll. 1886M96
| L. fraterna Oll. 1886M96
| L. marmorata (Arrow 1900) [=Oniscomorpha marmorata, Lepidopteryx marmorata]JG19
| L. seposita Oll. 1886M96
| L. turbata [incl. L. fasciculata]M96
|--L. cirrosa Pascoe 1860B14
`--+--L. decorataB14 [incl. L. ganydahensisM96]
`--L. moniliataB14
Trogossitidae incertae sedis:
PeltonyxaLB91
|--P. australis Blackb. 1891M96
`--P. pubescens Blackb. 1891M96
AncyronaLB91
|--A. aegra Oll. 1885M96
|--A. amica Oll. 1885M96
|--A. latebrosa Oll. 1885M96
|--A. laticeps Oll. 1885M96
`--A. vesca Oll. 1885M96
Latolaeva cassidioides Reitt. 1876M96
NeaspisM96
|--N. pusilla Blackb. 1891M96
`--N. variegata [=Soronia variegata; incl. N. subtrifasciata]M96
Peltoschema filicornis Reitt. 1880M96
NemosomiaL89b
|--N. elongatum [incl. N. corsicum]L89c
|--N. picta Léveillé 1889L89b
`--N. simoni Léveillé 1889L89c
Marnia Léveillé 1889L89d
|--*M. sipolisi Léveillé 1889L89d
`--M. sallei Léveillé 1890L90
LipaspisL89d
Melambia crenicollis [=Trogosita crenicollis]L89f
*Type species of generic name indicated
References
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[B74] Britton, E. B. 1974. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 62–89. Melbourne University Press.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.
[LC40] Laporte de Castelnau, F. 1840. Histoire Naturelle des Insectes Coléoptères vol. 2. P. Duménil: Paris.
[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).
[L02] Leschen, R. A. B. 2002. Trogossitidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 263–266. CRC Press.
[L89a] Léveillé, A. 1889a. Voyage de M. E. Simon au Venezuela (Décembre 1887–Avril 1888). Coléoptères, 3e mémoire. Temnochilides. Annales de la Société Entomologique de France, 6e série 9: 167–168.
[L89b] Léveillé, A. 1889b. Voyages de M. Émile Gounelle au Brésil. Temnochilides. Annales de la Société Entomologique de France, 6e série 9: 251–254.
[L89c] Léveillé, A. 1889c. Diagnose d’une espèce nouvelle de Nemosomia, et quelques observations synonymiques. Annales de la Société Entomologique de France, 6e série 9: viii–ix.
[L89d] Léveillé, A. 1889d. Diagnose d’un temnochilide nouveau, suivie de notes synonymiques sur le même groupe. Annales de la Société Entomologique de France, 6e série 9: xxi–xxii.
[L89e] Léveillé, A. 1889e. Observations synonymiques sur la famille des temnochilides. Annales de la Société Entomologique de France, 6e série 9: liv.
[L89f] Léveillé, A. 1889f. Rectifications suivantes relatives aux temnochilides (suite). Annales de la Société Entomologique de France, 6e série 9: lxii.
[L90] Léveillé, A. 1890. Description d’une nouvelle espèce de temnochilide. Annales de la Société Entomologique de France, 6e série 9: cxl.
[M96] Masters, G. 1896. Catalogue of the described Coleoptera of Australia. Supplement, part II. Dytiscidae, Gyrinidae, Hydrophyllidae, Staphylinidae, Pselaphidae, Paussidae, Silphidae, Scaphididae, Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydiidae, Cucujidae, Cryptophagidae, Lathridiidae, Mycetophagidae, Dermestidae, Byrrhidae, Parnidae, Heteroceridae. Proceedings of the Linnean Society of New South Wales 21 (Suppl.): 695–754.
[WH02] Worthy, T. H., & R. N. Holdaway. 2002. The Lost World of the Moa: Prehistoric life of New Zealand. Indiana University Press: Bloomington (Indiana).