Tyrina

Belongs within: Pselaphitae.

The Tyrina are a cosmopolitan group of pselaphine beetles with the third and fourth maxillary palpomeres elongate, typically the third palpomere being half as long as the fourth or longer, and the metacoxae dearly distant (Newton et al. 2001).

<==TyrinaNG05
    |--ParalasinusNG05
    |--Taiwanophodes Hlavác 2002NG05
    |--Nomuraius Hlavác 2002NG05
    |--Linan Hlavác 2002NG05
    |--Indophodes Hlavác 2002NG05
    |--Tyrodes clavatus Raffray 1895NG05
    |--Ryxabis Westwood 1870NG05, R04
    |    `--R. anthicoides Westwood 1870NG05
    |--Anitra Casey 1893NT01
    |    `--A. glaberula Casey 1893NT01
    |--Mipseltyrus Park 1953NT01
    |    |--M. leviniC90
    |    `--M. mirusNT01
    |--Labomimus Sharp 1883NG05, R04
    |    |--L. harmandi Raffray 1904R04
    |    |--L. heterocerus (Raffray 1882) [=Pselaphodes heterocerus; incl. P. antennatus Bryant 1915]NG05
    |    `--L. reitteri Sharp 1883R04
    |--Cedius LeConte 1849 [incl. Sinistrocedius Newton & Chandler 1989]NT01
    |    |--C. robustus Casey 1893R04
    |    |--C. spinosus Leconte 1850R04
    |    `--C. ziegleri Leconte 1850R04
    |--Pselaphodes Westwood 1870NG05, R04 [incl. Atherocolpus Raffray 1882R04]
    |    |--P. clavatus Raffray 1895NG05
    |    |--P. foveolatus Raffray 1892NG05
    |    |--P. heterocerus Raffray 1882R04
    |    |--P. simoni Raffray 1893R04
    |    |--P. sumatrensis Raffray 1896R04
    |    `--P. villosulus Westwood 1870NG05
    `--Tyrus Aubé 1833 [incl. Pseudotyrus Raffray 1890, Pytna Casey 1887]R04
         |--T. carinifer Casey 1897R04
         |--T. ‘clavatus’ Raffray 1895 non Raffray 1882R04
         |--T. compressicollis Schaufuss 1887R04
         |--T. consimilis Casey 1897R04
         |--T. corticalis (Raffray 1890)NG05 [=Pseudotyrus corticalisR04]
         |--T. corticinus (Casey 1887) [=Pytna corticina]R04
         |--T. histrio Schaufuss 1887R04
         |--T. humeralis (Aubé 1844) [=Hamotus humeralis; incl. T. compar]R04
         |--T. japonicus Sharp 1883R04
         |--T. mucronatus (see below for synonymy)R04
         |--T. myrmecophilus Bryant 1915NG05
         |--T. peyroniR04
         |--T. piceus King 1866R04
         `--T. semiruber Casey 1897R04

Tyrus mucronatus [incl. T. calcaratus Schaufuss 1887, T. costatus Fleischer 1888, T. insignis Reichenbach 1816, T. sanguineus, T. sexspinosus Schaufuss 1887]^R04

*Type species of generic name indicated

References

[C90] Chandler, D. S. 1990. Insecta: Coleoptera Pselaphidae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1175–1190. John Wiley & Sones: New York.

[NT01] Newton, A. F., M. K. Thayer, J. S. Ashe & D. S. Chandler. 2001. Staphylinidae Latreille, 1802. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 272–418. CRC Press: Boca Raton.

[NG05] Nomura, S., & I. Abd Ghani. 2005. Faunistic notes on the pselaphine species of the supertribes Goniaceritae, Pselaphitae and Clavigeritae from Malaysia and Singapore (Coleoptera: Staphylinidae: Pselaphinae). Serangga 10 (1–2): 1–36.

[R04] Raffray, A. 1904. Genera et catalogue des psélaphides [part III]. Annales de la Société Entomologique de France 73: 401–476, pls 1–3.