Belongs within: Anthracinae.
The Villini are a group of beeflies in which the pulvilli are reduced to minute tubercles (Lambkin et al. 2003). Members of the genus Villa have mostly hyaline wings (with at most the front margin infuscate) and often have brightly coloured stripes across the abdomen.
Characters (Lambkin et al. 2003): Antennae with apical stylomere present, basal stylomere rarely present, circlet of hairs absent; R2+3 arising from R with distinct basal 90° bend at i-r crossvein, squama fringed with scales; pulvilli indistinct, minute setose tubercles; laterotergite and mediotergites haired.
<==Villini [Villoestrini]
|--ExechohypopionLYG03
|--Cyananthrax cyanopterusGEEL09
|--Neodiplocampta paradoxaGEEL09
|--Paravilla separataGEEL09, KME08
|--DipaltaGEEL09
| |--D. banksiKME08
| `--D. serpentinaGEEL09
|--StonyxGEEL09
| |--S. cleliaH81
| `--S. clothoGEEL09
|--ChrysanthraxGEEL09
| |--C. cyprisGEEL09
| |--C. disparKME08
| `--C. edititiusGEEL09
|--LepidanthraxGEEL09
| |--L. campestrisH81
| |--L. inauratusBM76
| |--L. linguatusLYG03
| `--L. proboscideusE66
|--HemipenthesGEEL09
| |--H. catulinaKME08
| |--H. chimaera [=Villa (Hemipenthes) chimaera]E66
| |--H. jaennickeanaBT04
| |--H. morioKME08
| |--H. sinuosaKME08
| `--H. webberiKME08
|--PoecilanthraxGEEL09
| |--P. alyconKME08
| |--P. bicellataKME08 [=Exoprosopa bicellataLYG03]
| |--P. luciferGEEL09
| |--P. nigripennisKME08
| |--P. sackeniE66
| |--P. tegminipennisKME08
| `--P. willistoniiH81
`--VillaGEEL09
|--V. alternataKME08
|--V. arenicolaKME08
|--V. atraKME08
|--V. cyprisE66
|--V. faustinaE66
|--V. flavipilosaE66
|--V. flavocostalisE66
|--V. fulvianaKME08
|--V. fumicostaKME08
|--V. fuscicostataLYG03
|--V. gracilisKME08
|--V. handfordiKME08
|--V. harveyiE66
|--V. hypomelasKME08
|--V. johnsoniKME08
|--V. lateralisGEEL09
|--V. melasomaE66
|--V. molitorE66
|--V. nigraKME08
|--V. nigricaudaKME08 [=V. fulviana nigricaudaE66]
|--V. nigropectaKME08
|--V. pretiosaKME08
|--V. salebrosaE66
|--V. shawiiKME08
|--V. sinuosaE66
| |--V. s. sinuosaE66
| `--V. s. jaennickeanaE66
`--V. vestitaKME08
*Type species of generic name indicated
References
[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.
[BT04] Bybee, S. M., S. D. Taylor, C. R. Nelson & M. F. Whiting. 2004. A phylogeny of robber flies (Diptera: Asilidae) at the subfamilial level: molecular evidence. Molecular Phylogenetics and Evolution 30: 789–797.
[E66] Evans, H. E. 1966. The Comparative Ethology and Evolution of the Sand Wasps. Harvard University Press: Cambridge (Massachusetts).
[GEEL09] Greathead, D. J., N. L. Evenhuis & C. J. Einicker Lamas. 2009. Bombyliidae (bee flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 565–576. NRC Research Press: Ottawa.
[H81] Hall, J. C. 1981. Bombyliidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 589–602. Research Branch, Agriculture Canada.
[KME08] Kits, J. H., S. A. Marshall & N. L. Evenhuis. 2008. The bee flies (Diptera: Bombyliidae) of Ontario, with a key to the species of eastern Canada. Canadian Journal of Arthropod Identification 6: 1–52.
[LYG03] Lambkin, C. L., D. K. Yeates & D. J. Greathead. 2003. An evolutionary radiation of beeflies in semi-arid Australia: systematics of the Exoprosopini (Diptera: Bombyliidae). Invertebrate Systematics 17: 735–891.