Belongs within: Coleoptera.
Contains: Bostrichidae, Ptinidae, Dermestes, Megatominae.
The Bostrichoidea are a group of beetles predominantly adapted for dry habitats, united by modified cryptonephridism, aedeagal structure and the lack of a basal mandibular mola in the larva. Members include the Dermestidae whose larvae are scavengers on dry material of animal origin such as desiccated carcasses or shed hair or feathers (Lawrence & Britton 1991). Many dermestids may be recognised by their dense covering of setae or scales, and the presence of a median ocellus (Kingsolver 2002). The Orphilinae are of disputed affinities with proposed assignments to either Dermestidae or Nosodendridae (Ivie 2002a).
Endecatomus is a Holarctic genus of fungus-feeders that has been variously treated as its own family or included within the Bostrichidae. Species of Endecatomus have explanate pronotal margins bearing a row of projecting setae, and elytra bearing unisetose microtubercles arranged in an irregular, reticulate pattern (Ivie 2002b).
<==Bostrichoidea
|--+--Endecatomus Mellié 1847 [incl. Dictyalotus Redtenbacher 1847, Hendecatomus Bach 1852; Endecatomidae]I02b
| | |--E. dorsalis Mellié 1848B14
| | `--E. rugosus (Randall 1838)I02b
| `--+--BostrichidaeMW15
| `--PtinidaeMW15
`--Dermestidae [Dermestins, Dermestites, Megatoillidae]MW15
| i. s.: HexanodesAY04
| Thorictinae [Thorictidae, Thorictini]LB91
| |--Thorictodes Reitter 1875 [incl. Thaumaphrastus Blaisdell 1927]K02
| | `--T. heydeni Reitter 1875K02
| `--ThorictusA61
| |--T. capensisA61
| |--T. castaneus Germ. 1834G89
| |--T. germari Lucas 1846E12
| |--T. mauritanicus Lucas 1846E12
| `--T. puncticollis Lucas 1846E12
| TrinodinaeB14
| |--Trichelodes [Trichelodini]LB91
| | `--T. delicatulaLB91
| |--ThylodriniB14
| | |--TrichodriasB14
| | `--Thylodrias Motschulsky 1839 (see below for synonymy)K02
| | `--T. contractus Motschulsky 1839B14
| `--TrinodiniK02
| |--Trinodes hirtusLC40
| `--Apsectus LeConte 1854K02
| |--A. araneorum Beal 1959K02
| `--A. hispidus (Melsheimer 1844)K02
| Attageninae [Attagenini]LB91
| |--Novelsis Casey 1900 [incl. Genattus Sharp 1902, Paranovelsis Casey 1900]K02
| `--Attagenus Latreille 1802L02 [incl. Megatoma Kugelann 1792 non Herbst 1792K02]
| |--A. annulifer Laporte de Castelnau 1840LC40
| |--A. antennatus Laporte de Castelnau 1840LC40
| |--A. macellarius [=Dermestes macellarius]L02
| |--A. megatomaLC40
| |--A. pellioB70
| |--A. piceusB70
| |--A. posticalis Fairm. 1879G89
| |--A. schaefferiC01 [=Megatoma schaefferiLC40]
| |--A. trifasciatus [=Dermestes trifasciatus]L02
| |--A. undatusLC40
| |--A. unicolorOM91
| | |--A. u. unicolorOM91
| | `--A. u. japonicusOM91
| |--A. unifasciatus Fairm. 1860G89
| |--A. vagepictusF89
| `--A. vigintiguttatusLC40
| PhranodomaP92
| Globicornis rufitarsisLC40
|--+--DermestesMW15
| `--MegatominaeMW15
`--Orphilinae [Orphilini]MW15
|--OrphilodesB14
`--Orphilus Erichson 1846MW15, I02a
|--O. ater Erichson 1846I02a
|--O. dubius Wickham 1912I02a
|--O. niger (Rossi 1790)I02a
`--O. subnitidus LeConte 1861B14
Thylodrias Motschulsky 1839 [incl. Hospitopterus Pic 1921, Ignotus Slosson 1903, Thelydrias Gemminger & Harold 1868]K02
*Type species of generic name indicated
References
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[AY04] Austin, A. D., D. K. Yeates, G. Cassis, M. J. Fletcher, J. La Salle, J. F. Lawrence, P. B. McQuillan, L. A. Mound, D. J. Bickel, P. J. Gullan, D. F. Hales & G. S. Taylor. 2004. Insects ‘Down Under’—diversity, endemism and evolution of the Australian insect fauna: examples from select orders. Australian Journal of Entomology 43 (3): 216–234.
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.
[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.
[F89] Fairmaire, L. 1889. Coléoptères de l’intérieur de la Chine. 5e partie. Annales de la Société Entomologique de France, 6e série 9: 5–84.
[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.
[I02a] Ivie, M. A. 2002a. Nosodendridae Erichson 1846. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 224–227. CRC Press.
[I02b] Ivie, M. A. 2002b. Bostrichidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 233–244. CRC Press.
[K02] Kingsolver, J. M. 2002. Dermestidae Gyllenhal 1808. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 228–232. CRC Press.
[LC40] Laporte de Castelnau, F. 1840. Histoire Naturelle des Insectes Coléoptères vol. 2. P. Duménil: Paris.
[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.
[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).
[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.
[OM91] Okamoto, M., & K. Matsumoto. 1991. Observations on the behavior of the hypopus of Lardoglyphus konoi. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 555-559. SPB Academic Publishing: The Hague.
[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.