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Belongs within: Cucujoidea.
The Monotomidae are a group of elongate, parallel-sided beetles generally found under wood or in decaying vegetable matter (Bouchard 2014). Members of the Rhizophaginae have transverse fore coxae whereas the fore coxae are more or less rounded in Monotominae (Bousquet 2002).
Characters (from Lawrence & Britton 1991): Elongate, parallel-sided, subcylindrical to flattened, may be glabrous or clothed with decumbent setae, elytra may be truncate, exposing pygidium. Head usually abruptly narrowed posteriorly, forming distinct tempora (temples) behind eyes; antenna ten-segmented with one- or two-segmented club. Pronotum crenulate or dentate laterally; fore coxae globose, trochantins concealed, fore coxal cavities closed externally and internally. Mid coxae moderately separated, cavities open laterally. Tarsal formula 5-5-5 or 4-4-4, may be 5-5-4 in male. Abdomen with five visible ventrites, all free. Larva elongate, slightly flattened, slightly tapered posteriorly and lightly pigmented above but with tuberculate dorsal surface and sometimes lateral tergal processes on thorax and abdomen. Mala falciform; mandibular prostheca narrow and sometimes serrate; urogomphi weakly developed.
<==Monotomidae [Rhizophagidae]
| i. s.: Mimemodes laticeps (Macl. 1871)LB91, M96 [=Prostomis laticepsM96]
| MonotomopsisLB91
|--RhizophaginaeMW15
| |--Crowsonius meliponae Pakaluk & Ślipiński 1993B14
| `--Rhizophagus Herbst 1793B02
| | i. s.: R. aeneusC01
| | R. bipustulatus [=Lyctus bipustulatus]LC40
| | R. brunneusB02
| | R. parallelocollisB14
| | R. politus [=Lyctus politus; incl. Ips picipes]LC40
| | R. remotusMW15
| | R. unicolor Lucas 1846E12
| |--R. (Rhizophagus) [incl. Syringobidia Casey 1916]B02
| |--R. (Anomophagus Reitter 1907)B02
| `--R. (Eurhizophagus Méquignon 1909)B02
`--Monotominae [Lenacinae, Thioninae]MW15
|--Lenax [Lenacini]B02
| `--L. mirandus Sharp 1877B14
|--ThioniniB02
| |--Thione Sharp 1899B02
| | `--T. championiB02
| `--Shoguna Lewis 1889B02, L90
| |--S. striata Arrow 1900JG19
| `--S. termitiformis (Fairmaire 1883) [incl. S. polita Arrow 1900]JG19
|--Monotoma Herbst 1793 [Monotomini]B02
| | i. s.: M. americanaLB91
| | M. longicollisB02
| | M. picipesB02 [=Lyctus picipesLC40]
| | M. rufaM96
| |--M. (Gyrocecis Thomson 1863)B02
| | `--M. (G.) myrmecophila Bousquet & Laplante 2000B02
| `--M. (Monotomina Nikitsky 1986)B02
| `--M. (M.) quadrifoveolata Aubé 1837B02
`--EuropiniB02
|--Bactridium LeConte 1861 non Kunze 1817 (ICBN)B02
|--Aneurops Sharp 1900B02
|--Pycnotomina Casey 1916B02
| `--P. cavicolle (Horn 1879)B02
|--Macreurops Casey 1916B02
| `--M. longicollis (Horn 1879)B02
|--Phyconomus LeConte 1861B02
| `--P. marinus (LeConte 1858)B02
|--Leptipsius Casey 1916B02
| |--L. dilutus Casey 1916B02
| `--L. striatus (LeConte 1858)B02
|--Hesperobaenus LeConte 1861B02
| |--H. apicalis Reitt. 1872FS90
| `--H. subtestaceus [=Phyconomus subtestaceus]B02
|--Europs Wollaston 1854 [incl. Nomophloeus LeConte 1861]B02
| |--E. depressusRS72
| `--E. luridipennis [=Phyconomus luridipennis]B02
`--MimemaB02
|--M. pallidum [=Phyconomus pallidus]B02
`--M. tricolor [=Phyconomus tricolor]B02
*Type species of generic name indicated
References
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[B02] Bousquet, Y. 2002. Monotomidae Laporte 1840. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 319–321. CRC Press.
[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.
[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.
[LC40] Laporte de Castelnau, F. 1840. Histoire Naturelle des Insectes Coléoptères vol. 2. P. Duménil: Paris.
[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).
[L90] Léveillé, A. 1890. Rectification synonymique. Annales de la Société Entomologique de France, 6e série 9: ccxi.
[M96] Masters, G. 1896. Catalogue of the described Coleoptera of Australia. Supplement, part II. Dytiscidae, Gyrinidae, Hydrophyllidae, Staphylinidae, Pselaphidae, Paussidae, Silphidae, Scaphididae, Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydiidae, Cucujidae, Cryptophagidae, Lathridiidae, Mycetophagidae, Dermestidae, Byrrhidae, Parnidae, Heteroceridae. Proceedings of the Linnean Society of New South Wales 21 (Suppl.): 695–754.
[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.
[RS72] Rathore, Y. S., & C. S. Sengar. 1972. New records of nitidulid and rhizophagid beetles on maize cobs in the U.P. Tarai. Journal of the Bombay Natural History Society 69 (1): 208–209.