Belongs within: Pseudocaecilioidea.
The Pseudocaeciliinae are a group of barklice with two-segmented tarsi, known predominantly from south-east Asia and the Pacific (New & Lienhard 2007).
<==Pseudocaeciliinae [Meplerinae, Pseudoscottiinae]NL07
|--*Allopsocus marginalis Banks 1920NL07
|--ScottiellaNL07
|--Platyocaecilius Li 2002L02b
| `--*P. parallelivenius Li 2002L02b
|--Fashengocaecilius Lienhard 2003L03 [=Calocaecilius Li 2002 non Mockford 1974L02b]
| `--*F. biaduncus (Li 1992) [=Allocaecilius biaduncus, *Calocaecilius biaduncus]L03
|--Phyllocaecilius Li 2002L02b
| `--*P. atrichus Li 2002L02b
|--Chorocaecilius Li 2000L02a
| |--C. brunellus (Tillyard 1923) (see below for synonymy)SN08
| `--C. decemdentatus Li 2000L02a
|--Phallocaecilius Lee & Thornton 1967L02b
| |--*P. hirsutus (Thornton 1961) [=Pseudocaecilius hirsutus]NL07
| `--P. sentosus Li 1999L02a
|--Thelocaecilius Li 2002L02b
| |--*T. mecokeratus Li 2002L02b
| `--T. papillatus (Li 1995) [=Heterocaecilius papillatus]L02b
|--Neocaecilius Li 2000L02a
| |--*N. campanula [=Heterocaecilius campanula]NL07
| |--N. granulatus (Lee & New 1992)NL07
| |--N. mutatus (Lee & New 1992)NL07
| `--N. qianshanensis Li 2000L02a
|--Mesocaecilius Okamoto 1910NL07, L02a [Mesocaeciliinae]
| |--*M. quadrimaculatus Okamoto 1910L02a
| |--M. bellus Li 2002L02b
| |--M. decorosus Li 2002L02b
| |--M. elegans Li 1997L02b
| |--M. euryopterus Li 2002L02b
| |--M. lepidus Li 2002L02b
| `--M. paraornatus (New 1977) [=Pseudocaecilius paraornatus]L02b
|--Ophiodopelma Enderlein 1908L02a
| |--*O. ornatipenne Enderlein 1908NL07, L02a
| |--O. anocella Li 2002L02b
| |--O. fluctosa Li 1995L02a
| |--O. glyptocephalum Lienhard 1985NL07
| |--O. hieroglyphicum (Enderlein 1903)NL07
| |--O. multipunctatum (Hagen 1859)NL07
| |--O. permaculatum Lee & Thornton 1967NL07
| |--O. pictipenna Lee & Thornton 1967NL07
| |--O. polyspila Li 1993L02b
| `--O. semiceps Lee & Thornton 1967NL07
|--Levucaecilius Smithers 1995NL07, S95
| |--*L. greenwoodi [=Pseudocaecilius greenwoodi, Orbocaecilius greenwoodi]NL07
| |--L. albicrus (Thornton 1981) [=Heterocaecilius albicrus]S95
| |--L. apicalis (Thornton 1981) [=Heterocaecilius apicalis]S95
| |--L. dardanus (Lee & Thornton 1967)S95 [=Heterocaecilius dardanusNL07, Neocaecilius dardanusNL07]
| |--L. hilli Smithers 1995NL07
| |--L. panicus (Lee & Thornton 1967) [=Heterocaecilius panicus]S95
| |--L. pictus (Thornton 1981) [=Heterocaecilius pictus]S95
| |--L. simplex (Lee & Thornton 1967)NL07, L02a [=Heterocaecilius simplexNL07]
| |--L. veitchi [=Heterocaecilius veitchi]S95
| `--L. volatus (Lee & Thornton 1967) [=Heterocaecilius volatus]S95
|--Lobocaecilius Lee & Thornton 1967L02a
| |--*L. cynara Lee & Thornton 1967L02a
| |--L. arcuatus Vaughan, Thornton & New 1989NL07
| |--L. bifurcus Li 1993L02a
| |--L. drepanus New & Thornton 1976NL07
| |--L. ligur Lee & New 1992NL07
| |--L. longifurca Lee & Thornton 1967NL07
| |--L. pahangensis Lee & New 1992NL07
| |--L. perductivirgus Li 2002L02b
| |--L. pilosus Lee & New 1992NL07
| |--L. quadripartitus Li 1995L02b
| |--L. thorntoni Lee & New 1992NL07
| `--L. viridis Lee & New 1992NL07
|--Allocaecilius Lee & Thornton 1967L02a
| |--*A. heterothrax Lee & Thornton 1967L02a
| |--A. albidorsualis Li 1993L02a
| |--A. arcuatus Li 2002L02b
| |--A. elongatus Lee & Thornton 1967NL07
| |--A. fusciceps Lee & Thornton 1967NL07
| |--A. gombakensis Lee & New 1992NL07
| |--A. guangdongicus Li 1993L02a
| |--A. maculatus Lee & Thornton 1967NL07
| |--A. selangorensis New & Thornton 1975NL07
| |--A. sinensis Lee & Thornton 1967L02a
| |--A. smithersi Lee & New 1992NL07
| `--A. tenuilongus Li 1995L02a
|--Mepleres Enderlein 1926SN08 [incl. Meniscopsocus Li 1993L02b, Pseudoscottiella Badonnel 1946L02b]
| |--*M. maeandricus Enderlein 1926SN08
| |--M. cornutus (Lee & New 1992)NL07 [=Pseudoscottiella cornutaL02b]
| |--‘Pseudoscottiella’ decolor Badonnel 1955B55
| |--‘Pseudoscottiella’ hyalina Badonnel 1955B55
| |--‘Pseudoscottiella’ immaculata Badonnel 1955B55
| |--M. indicus (Lee & New 1992)NL07
| |--M. limbatus (Enderlein 1908)L02a
| |--M. longicellus Li 2002L02b
| |--M. longitudinalis Li 2002 [=M. longitudianlis]L03
| |--‘Pseudoscottiella’ maculata Broadhead & Richard 1982L02b
| |--‘*Pseudoscottiella’ megopsNL07
| |--M. nigroguttatus (Karny 1925)NL07 [=Pseudoscottiella nigraguttataL02b]
| |--M. ocellatus Li 2002L02b
| |--M. oresbius Li 2002L02b
| |--M. parvicellus Li 2002L02b
| |--M. pisiformis (Lee & New 1992)NL07
| |--M. plenimacularis Li 2002L02b [=Pseudoscottiella plenimacularisL02a]
| |--M. proboscideus Li 2002L02b
| |--M. procurrens Li 2002L02b
| |--M. radialis (Thornton 1984)NL07
| |--M. raphalis (Lee & New 1992)NL07
| |--M. rotundatus (New 1974) [=Pseudoscottiella rotundata, Meniscopsocus rotundatus]SN08
| |--M. similis (Lee & New 1992)NL07
| |--M. sinicus (Li 1993)L02b [=*Meniscopsocus sinicusNL07]
| |--M. solenocerus (Lee & New 1992)NL07
| |--M. suella (Thornton 1984)NL07 [=Pseudoscottiella suellaL02b]
| |--M. tanei (Smithers 1977) [=Pseudoscottiella tanei, Meniscopsocus tanei]SN08
| |--M. transversus (Banks 1937)NL07
| |--‘Pseudoscottiella’ tuberculata Badonnel 1955B55
| |--M. ulterior Li 2002L02b
| |--M. unitus Li 2002L02b
| |--M. urbanus (Vaughan, Thornton & New 1991)NL07
| |--M. vilus (Lee & New 1992)NL07
| |--‘Pseudoscottiella’ yeni New 1974L02b
| `--M. yunnanicus Li 2002L02b
|--Pseudocaecilius Enderlein 1903Y02 [incl. Hageniella Enderlein 1903 (preoc.)L02a]
| |--P. citricola (Ashmead 1879)NL07 (see below for synonymy)
| |--P. angustus (Enderlein 1903)NL07
| |--P. bibulbus Li 1993L03, L02a [=P. bibalbus (l. c.)L03]
| |--P. bicostatus Li 2002L02b
| |--P. ceratocercus Li 2002L02b
| |--P. claggi (Banks 1937)NL07
| |--P. clunialis Vaughan, Thornton & Neo 1991NL07
| |--P. cornutus Lee & Thornton 1967NL07
| |--P. danaus Lee & New 1992NL07
| |--P. euryocercus Li 2002 [=P. eurocerus]L03
| |--P. formosanus (Banks 1937)L02a
| |--P. galactozonalis Li 2002 [=P. galactozonacis]L03
| |--P. helicoides Lee & Thornton 1967NL07
| |--P. immaculatus Li 2002L02b
| |--P. inaequalis (Banks 1916)NL07
| |--P. innotatus Banks 1937NL07
| |--P. kagoshimensisY02
| |--P. largicellus Li 2002L02b
| |--P. machadoi Badonnel 1955B55
| |--P. monotaeniatus Li 2002 [=P. monotaenitus, P. monotaenius]L03
| |--P. nitoris (Banks 1937)NL07
| |--P. octomaculatus (Li 1995) [=Heterocaecilius octomaculatus]L02b
| |--P. otiosus (Banks 1937)NL07
| |--P. papillaris Li 2002L02b
| |--P. plagiozonalis Li 1995L02a
| |--P. productus New & Thornton 1976NL07
| |--P. pusillus (Banks 1931)NL07
| |--P. ranus Thornton 1984NL07
| |--P. serratus Lee & New 1992NL07
| |--P. setifer Vaughan, Thornton & New 1991NL07
| |--P. sexdentatus Li 2002L02b
| |--P. tahitiensis (Karny 1926) [=Epipsocus tahitiensis]M93
| |--P. tenellus Enderlein 1926NL07
| |--P. testaceus Enderlein 1903NL07
| |--P. undecimimaculatus Li 1995L02b
| |--P. utricularis Li 1999L02a
| |--P. venimaculatus Li 2002L02b
| |--P. wellsae Lee & New 1992NL07
| `--‘Psocus’ zonatus [=*Hageniella zonata]NL07
`--Heterocaecilius Lee & Thornton 1967L02a [incl. Orbocaecilius Li 1999L03, Obrocaecilius (l. c.)L03]
|--*H. minutus Lee & Thornton 1967SN08 [=H. minotusNL07]
|--H. abacutidens Li 2002L02b
|--H. analis Vaughan, Thornton & New 1991NL07
|--H. anareolatus New 1993NL07
|--H. anomalus (Thornton 1961)Y02, L02b [=Orbocaecilius anomolusL02b]
|--H. argutus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) argutus]L03
|--H. bicruris (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) bicruris]L03
|--H. bidigitatus Li 1992L02a [=Orbocaecilius bidigitatusL03]
|--H. bifarius (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) bifarius]L03
|--H. brachystigmus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) brachystigmus]L03
|--H. brevifurca Thornton 1984NL07
|--H. challengerius Lee & New 1992NL07
|--H. circulicellus Li 1995L02b [=Orbocaecilius circulicellusL03]
|--H. dasoceratus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) dasoceratus]L03
|--H. deloneurous Li 2002L02b
|--H. dentalis Li 2002L02b
|--H. duodecidentus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) duodecidentus]L03
|--H. fasciatus Lee & Thornton 1967NL07
|--H. flaviformis Lee & New 1992NL07
|--H. fumosus Thornton 1984NL07
|--H. fuscipalpus Lee & Thornton 1967NL07
|--H. fuscusY02
|--H. gressitti Lee & Thornton 1967NL07
|--H. hepaticivenus Li 2002L02b
|--H. hoplites Lee & New 1992NL07
|--H. huananensis Li 2002L02b
|--H. huaxiensis Li 2002L02b
|--H. imparilis (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) imparilis]L03
|--H. jinghongicus Li 1995L02a
|--H. kobus Lee & Thornton 1967NL07
|--H. lachlani (Enderlein 1903) [=Pseudocaecilius lachlani]SN08
|--H. leei New 1993NL07
|--H. liangi Li 1991L02b
|--H. loiaoi New 1991L02a
|--H. longicornis (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) longicornis]L03
|--H. longipenna Lee & Thornton 1967NL07
|--H. longispinus Lee & Thorton 1967NL07
|--H. maculans Li 1991L02b
|--H. maculifornis (Thornton 1961)L02b [=Pseudocaecilius maculifronsL02a]
|--H. maculosus Lee & New 1992NL07
|--H. marginatus Lee & New 1992NL07
|--H. mecotrichus Li 2002L02b
|--H. odontothelus Li 2002L02b
|--H. orientalis Lee & New 1992NL07
|--H. ornatus (Enderlein 1903) [=Pseudocaecilius ornatus]NL07
|--H. papillosus Vaughan, Thornton & New 1991NL07
|--H. paulicellus (Li 1999)NL07, L02a [=*Orbocaecilius paulicellusL02a]
|--H. platokeratis Li 1993L02a
|--H. pseudoanomalus Li 1993L02b [=Obrocaecilius (l. c. for Orbocaecilius) pseudoanomalusL03]
|--H. quadrangulus Li 1992L02b
|--H. reticulatus Lee & New 1992NL07
|--H. rhadnus Thornton 1984NL07
|--H. sedecimidentus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) sedecimidentus]L03
|--H. stenokeratis Li 1993L02a
|--H. stiliger Lee & Thornton 1967NL07
|--H. taipingensis Garcia Aldrete 1999NL07
|--H. tartareus Li 2002L02b
|--H. vulturius (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) vulturius]L03
`--H. xihuicus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) xihuicus]L03
Chorocaecilius brunellus (Tillyard 1923) [=Caecilius brunellus, Heterocaecilius brunellus, Pseudocaecilius brunellus; incl. H. diogenes Lee & Thornton 1967, *Chorocaecilius diogenes]SN08
Pseudocaecilius citricola (Ashmead 1879)NL07 [=Psocus citricolaM93; incl. Pse. elutus var. africanus Badonnel 1931L02a, Elipsocus criniger Perkins 1899M93, Kilauella crinigerM93, Pseudocaecilius crinigerM93, *Pse. elutus Enderlein 1903NL07, L02b, Trichopsocus (*Pseudocaecilius) elutusL02a, Trichopsocus indicatus Navás 1934M93, Pse. medianicostatus Li 1993L02a, Caecilius pretiosus Banks 1920M93, Pse. pretiosusM93, Pse. wolcotti Banks 1924M93]
*Type species of generic name indicated
References
[B55] Badonnel, A. 1955. Psocoptères de l’Angola. Diamang Publicações Culturais 26: 11–267.
[L02a] Li F. 2002a. Psocoptera of China vol. 1. Science Press: Beijing.
[L02b] Li F. 2002b. Psocoptera of China vol. 2. Science Press: Beijing.
[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.
[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.
[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.
[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.
[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529–561.
[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371–400.