Belongs within: Scincomorpha.
Contains: Eremiascincus, Chalcides, Lygosoma (Rhodona), Egernia, Liopholis, Tiliqua, Cyclodomorphus, Ctenotus, Hemiergis, Riopa, Dasia, Mochlus, Nodorha, Norbea, Ophioscinus, Otosaurus, Saiphos, Tropidophorus, Lerista, Mabuya, Sphenomorphus, Scincella, Ablepharus, Lipinia, Panaspis, Lampropholis, Carlia, Emoia, Cryptoblepharus, Morethia, Leiolopisma, Oligosoma, Glaphyromorphus, Lygisaurus, Saproscincus, Scincinae.
The Scincidae, skinks, are the largest of the traditionally-recognised families of lizards, with more than 1400 species. Many skinks have smooth, glass-like scales (though there are notable exceptions), and many are elongate. Possible characters supporting the skinks as a group are the structure of the osteoderms and the present of a secondary palate (Rieppel 1988). Monophyly of the family has been challenged in the past, with the Feyliniinae in particular being on occasion treated separately, but the presence of scroll-like palatines may support a clade including the Feyliniinae and Lygosominae (Rieppel 1988).
Scincidae
|--Acontias [Acontiadae, Acontianinae, Acontiidae, Acontinae]MH11
| `--A. meleagrisMH11
`--+--ScincinaeMH11
`--+--Feyliniinae [Feyliniidae]MH11
| |--Voeltzkowia fierinensisU57, MH11
| `--FeyliniaBH10
| |--F. curroriBH10
| `--F. polylepisMH11
`--ScelotinaeMH11
|--Proscelotes eggeliMH11
`--ScelotesMH11
|--S. bipesMH11
|--S. kafferMH11
|--S. kasneriMH11
|--S. gronoviMH11
|--S. mirusMH11
`--S. sexlineatusMH11
Scincidae incertae sedis:
EremiascincusC18
Trachysaurus rugosusG60a
Macroscincus coctei Duméril & Bibron 1839H81
Gongylomorphus bojeriiH81
|--G. b. bojeriiH81
`--G. b. borbonica Vinson 1969H81
ChalcidesR88
Plestiodon obsoletusBH10
TyphlacontiasU57
LygosominaeH11
| i. s.: Lygosoma Hardwich & Gray 1827C18
| | i. s.: L. aeneumHD09
| | L. assataDD61
| | |--L. a. assataDD61
| | `--L. a. tayloriDD61
| | L. bowringii (Günther 1864)C18
| | L. cophia Boulenger 1908M52
| | L. darlingtoni Loveridge 1933M52
| | L. dendyiHD09
| | L. grandeHD09
| | L. lineoocellatumHD09
| | L. luzonense Boulenger 1895M52
| | L. mulleri Peters 1878R13, LF96
| | L. ornatumHD09
| | L. pumila Boulenger 1887M52
| | L. punctulata Peters 1871M52
| | L. relictumB28
| | L. scutirostra Peters 1873M52
| | L. smithiiHD09
| |--L. (Lygosoma) [incl. Podophis Wiegmann 1834]M52
| | |--L. (L.) larutense Boulenger 1900B00
| | |--L. (L.) quadripes (Linné 1766) (see below for synonymy)M52
| | `--L. (L.) sumatrenseB00
| |--L. (Omolepida)G60b
| | |--L. (O.) australe Gray 1839G60b, M52
| | |--L. (‘Homolepida’) casuarinae (see below for synonymy)LF96
| | `--L. (O.) deplanchei [incl. Hinulia tetragonurus]R13
| |--L. (Rhodona)G60b
| `--L. (Siaphops) gracile [=L. gracilis; incl. Mocoa micropus]R13
| EgerniaH11
| LiopholisC18
| Bellatorias Wells & Wellington 1984H11, C18
| |--B. frerei (Günther 1897)C18
| |--B. major (Gray 1845)C18
| `--B. obiri (Wells & Wellington 1985)C18
| Lissolepis Peters 1872H11, C18
| |--L. coventryi (Storr 1978)C18
| `--L. luctuosa iPeters 1866)C18
| TiliquaH11
| CyclodomorphusH11
| CtenotusG74
| HemiergisG60b
| RiopaM52
| Ateuchosaurus Gray 1845 [incl. Lygosaurus Hallowell 1860]M52
| |--*A. chinensis Gray 1845M52
| |--A. browni Van Denburgh 1912M52
| |--A. pellopleurus (Hallowell 1860) [=*Lygosaurus pellopleurus]M52
| |--A. salsburyi Schmidt 1927M52
| `--A. sowerbyi Stejneger 1924M52
| DasiaM52
| Eumecia Bocage 1870M52
| |--*E. anchietae Bocage 1870M52
| `--E. johnstoni Boulenger 1897M52
| MochlusM52
| NodorhaM52
| NorbeaM52
| OphioscincusM52
| OtosaurusM52
| Ristella Gray 1839M52
| |--*R. rurkii Gray 1839M52
| |--R. beddomii Boulenger 1887M52
| |--R. guentheri Boulenger 1887M52
| `--R. travancorica Beddome 1870M52
| SaiphosM52
| Squamicilia Mittleman 1952M52
| |--*S. isodactyla (Günther 1864) [=Eumeces isodactylus]M52
| `--S. producta Boulenger 1909M52
| Tribolonotus Duméril & Bibron 1839 [incl. Pediporus Roux 1930]M52
| |--*T. novaeguineae (Schlegel 1834) [=Zonurus novaeguineae]M52
| |--T. blanchardi Burt 1930M52
| |--T. gracilis de Rooij 1909M52
| `--T. schmidti Burt 1930 [=*Pediporus schmidti]M52
| TropidophorusM52
| ‘Monophorus’ Cope 1892 (n. d.) non Grillo 1877M52
| ‘Oncopus’ Cope 1892 (n. d.) nec Herrich-Schaeffer 1855 nec Thorell 1876 nec Karsten 1882 (ICBN)M52
| Ollochirus Cope 1892 (n. d.)M52
| Tridentulus Cope 1892 (n. d.)M52
| Furcillus Cope 1892 (n. d.)M52
| Haploscincus Cope 1892 (n. d.)M52
| Dimeropus Cope 1892 (n. d.)M52
| LeristaM52
|--MabuyaG74
|--+--SphenomorphusG74
| `--+--ScincellaG74
| |--AblepharusG74
| `--+--Lobulia Greer 1974G74
| | |--*L. elegans (Boulenger 1897) [=Lygosoma elegans]G74
| | |--L. brongersmai Zweifel 1972G74
| | |--L. morokana Parker 1936G74
| | |--L. phaeodes Vogt 1932G74
| | `--L. stanleyana Boulenger 1897G74
| `--+--LipiniaG74
| `--Prasinohaema Greer 1974G74
| |--*P. flavipes (Parker 1936) [=Lygosoma flavipes]G74
| |--P. parkeri Smith 1937G74
| |--P. prehensicauda Loveridge 1945G74
| |--P. semoni Oudemans 1894G74
| `--P. virens Peters 1881G74
| |--P. v. virensG74
| `--P. v. anolisG74
`--+--+--+--PanaspisG74
| | |--Cophoscincopus Mertens 1934 [=Cophoscincus Vaillant 1884 non Peters 1867]G74
| | | `--C. durus (Cope 1862) (see below for synonymy)G74
| | `--Afroablepharus Greer 1974G74
| | |--*A. wahlbergi (Smith 1849) [=Cryptoblepharus wahlbergi]G74
| | |--A. duruarus Monard 1949G74
| | |--A. megalurus Nieden 1913G74
| | |--A. seydeli de Witte 1933G74
| | |--A. tancredi Boulenger 1909G74
| | `--A. wilsoni Werner 1914G74
| `--+--LampropholisG74
| |--Geomyersia Greer & Parker 1968G74
| | `--*G. glabra Greer & Parker 1968G74
| |--Notoscincus Fuhn 1969G74
| | |--*N. wotjulum (Glauert 1959)G74 [=Ablepharus wotjulumG74, N. ornatus wotjulumPP13]
| | |--N. butleri Storr 1979C18
| | `--N. ornatus (Broom 1896)HD04, G74 [=Ablepharus ornatusG74]
| `--+--CarliaG74
| `--Menetia Gray 1845G74
| |--*M. greyii Gray 1845G74 [=Ablepharus greyiiM52; incl. M. amaura Storr 1978C18]
| |--M. alanae Rankin 1979C18
| |--M. concinna Sadlier 1984C18
| |--M. maini Storr 1976C18
| `--M. surda Storr 1976C18
| |--M. s. surdaADD08
| `--M. s. cresswelliADD08
`--+--+--Tachygia Mittleman 1952 [=Liosoma Fitzinger 1843 non Brandt 1834]G74
| | |--*T. microlepis (Duméril & Bibron 1839)G74 (see below for synonymy)
| | `--T. garnieri Bavay 1869M52
| |--Phoboscincus Greer 1974G74
| | |--*P. bocourti (Brocchi 1876)G74 [=Eumeces bocourtiG74, Lygosoma (Riopa) bocourtiR13]
| | `--P. garnieri Bavay 1869G74
| `--Eugongylus Fitzinger 1843G74
| |--E. rufescens (Shaw 1802)G74 (see below for synonymy)
| |--E. albofasciolatus Günther 1872G74
| |--E. boettgeri Sternfeld 1918M52
| |--E. mentovarius Boettger 1895M52
| `--E. sulaensis Kopstein 1927G74
`--+--+--EmoiaG74
| `--+--CryptoblepharusG74
| `--MorethiaG74
`--+--LeiolopismaG74
|--OligosomaG74
|--Proablepharus Fuhn 1969G74, C18
| |--*P. reginae (Glauert 1960) [=Ablepharus reginae]G74
| |--P. broomensis Lönnberg & Andersson 1913G74
| |--P. davisi Copland 1952G74
| `--P. tenuis Broom 1896G74
|--Anotis Bavay 1869 [incl. Nannoscincus Günther 1872]G74
| |--*A. mariae Bavay 1869G74 (see below for synonymy)
| |--A. gracilis Bavay 1869G74
| |--A. graciloides Lönnberg & Andersson 1913G74
| |--A. initialis Werner 1910M52
| |--A. maccoyi Lucas & Frost 1894G74
| |--A. scarffi Boulenger 1915M52
| `--A. slevini Loveridge 1941G74
`--Cyclodina Girard 1857WH02, G74
|--*C. aenea Girard 1857 [=Leiolopisma aeneum]G74
|--C. alani (Robb 1970)WH02
|--C. macgregori (Robb 1975)WH02
|--C. northlandi Worthy 1991WH02
|--C. oliveri (McCann 1955)WH02
|--C. ornata (Gray 1843)WH02
`--C. whitakeri Hardy 1977WH02
Asymblepharus sikimmensisAS09
Euprepis auratusAS09
TyphlosaurusAS09
|--T. aurantiacusR93
|--T. cregoiR93
|--T. lineatusR93
|--T. lomiiAS09
`--T. vermisBP87
Sauriscus cooki Estes 1964B93
Acritoscincus Wells & Wellington 1985 [incl. Bassiana]C18
|--A. duperreyi (Gray 1838)C18
|--A. platynotus (Peters 1881)C18
`--A. trilineatus (Gray 1838)C18
Anespischetosia Wells & Wellington 1985C18
`--A. maccoyi (Lucas & Frost 1894)C18
Anomalopus Duméril & Duméril 1851C18
|--*A. verreauxii (Duméril & Duméril 1851)M52, C18
|--A. brevicollis Greer & Cogger 1985C18
|--A. gowi Greer & Cogger 1985C18
|--A. leuckartii (Weinland 1862)C18
|--A. mackayi Greer & Cogger 1985C18
|--A. pluto Ingram 1977C18
`--A. swansoni Greer & Cogger 1985C18
Calyptotis De Vis 1886C18
|--C. lepidorostrum Greer 1983C18
|--C. ruficauda Greer 1983C18
|--C. scutirostrum (Peters 1873)C18
|--C. temporalis Greer 1983C18
`--C. thorntonensis Greer 1983C18
Carinascincus Wells & Wellington 1985C18
|--C. coventryi (Rawlinson 1975)C18
|--C. greeni (Rawlinson 1975)C18
|--C. metallicus (O’Shaughnessy 1874)C18
|--C. microlepidotus (O’Shaughnessy 1874)C18
|--C. ocellatus (Gray 1845)C18
|--C. orocryptus (Hutchinson, Schwaner & Medlock 1988)C18
|--C. palfreymani (Rawlinson 1974)C18
`--C. pretiosus (O’Shaughnessy 1874)C18
Coeranoscincus Wells & Wellington 1984C18
|--C. frontalis (De Vis 1888)C18
`--C. reticulatus (Günther 1873)C18
Coggeria Couper, Covacevich et al. 1996C18
`--C. naufragus Couper, Covacevich et al. 1996C18
Concinnia Wells & Wellington 1984C18
|--C. amplus (Covacevich & McDonald 1980)C18
|--C. brachysoma (Lönnberg & Andersson 1915)C18
|--C. frerei (Greer 1992)C18
|--C. martini Wells & Wellington 1985C18
|--C. sokosoma (Greer 1992)C18
|--C. tenuis (Gray 1831)C18
`--C. tigrina (De Vis 1888)C18
Eroticoscincus Wells & Wellington 1984C18
`--E. graciloides (Lönnberg & Andersson 1913)C18
Eulamprus Fitzinger 1843C18 [=Hinulia Gray 1845M52]
|--*E. quoyii (Duméril & Bibron 1839)M52, C18 [=Lygosoma quoyiM52, *Hinulia quoyiM52]
|--E. heatwolei Wells & Wellington 1984C18
|--E. kosciuskoi (Kinghorn 1932)C18
|--E. leuraensis Wells & Wellington 1984C18
`--E. tympanum (Lönnberg & Andersson 1913)C18
|--E. t. tympanumC18
`--E. t. marniaeC18
GlaphyromorphusC18
Gnypetoscincus Wells & Wellington 1984C18
`--G. queenslandiae (De Vis 1890)C18
Harrisoniascincus Wells & Wellington 1985C18
`--H. zia (Ingram & Ehmann 1981)C18
Liburnascincus Wells & Wellington 1984C18
|--L. artemis Hoskin & Couper 2015C18
|--L. coensis (Mitchell 1953)C18
|--L. mundivensis (Broome 1898)C18
`--L. scirtetis (Ingram & Covacevich 1980)C18
LygisaurusC18
Nangura Covacevich, Couper & James 1993C18
`--N. spinosa Covacevich, Couper & James 1993C18
Pseudemoia Fuhn 1967C18
|--*P. spenceri (Lucas & Frost 1894) [=Emoa spenceri, Leiolopisma spenceri]G74
|--P. baudini (Greer 1982)C18
|--P. cryodroma Hutchinson & Donnellan 1992C18
|--P. entrecasteauxii (Duméril & Bibron 1839)C18
|--P. pagenstecheri (Lindholm 1901)C18
`--P. rawlinsoni (Hutchinson & Donnellan 1988)C18
SaproscincusC18
Techmarscincus Wells & Wellington 1985C18
`--T. jigurru (Covacevich 1984)C18
Austroablepharus Couper, Hoskin et al. 2018C18
|--A. barrylyoni (Couper, Limpus et al. 2010) [=Proablepharus barrylyoni]C18
|--A. kinghorni (Copland 1947) [=Proablepharus kinghorni]G74
`--A. naranjicaudus (Greer, Fisher & Horner 2004) [=Proablepharus naranjicaudus]C18
Pygmaeascincus Couper & Hoskin 2014C18
|--P. koshlandae (Greer 1991) [=Menetia koshlandae]C18
|--P. sadlieri (Greer 1991) [=Menetia sadlieri]C18
`--P. timlowi (Ingram 1977) [=Menetia timlowi]C18
Karma Wells 2009 [incl. Silvascincus Skinner, Hutchinson & Lee 2013]C18
|--K. murrayi (Boulenger 1887) [=Concinnia murrayi, Silvascincus murrayi]C18
`--K. tryoni (Longman 1918) [=Concinnia tryoni, Silvascincus tryoni]C18
Magmellia Wells 2009 [incl. Tumbunascincus Skinner, Hutchinson & Lee 2013]C18
`--*M. luteilateralis (Covacevich & McDonald 1980) (see below for synonymy)C18*Anotis mariae Bavay 1869G74 [=Lygosoma (Siaphops) marieiR13; incl. *Nannoscincus fuscus Günther 1872G74]
Cophoscincopus durus (Cope 1862) [=Tiliqua durus; incl. *Cophoscincus simulans Vaillant 1884, *Cophoscincopus simulans]G74
Eugongylus rufescens (Shaw 1802)G74 [=Lacerta rufescensM52; incl. Eumeces oppelii Duméril & Bibron 1839G74, *Eugongylus oppeliiG74]
Lygosoma (‘Homolepida’) casuarinae [=Cyclodus casuarinae, Omolepidota casuarinae; incl. Homolepida nigricans Peters 1874, Hemisphaeriodon tasmanicum Lucas & Frost 1893]LF96
Lygosoma (Lygosoma) quadripes (Linné 1766) [=Anguis quadrupes, *Podophis quadrupes; incl. Lacerta serpens, *Lygosoma serpens]M52
*Magmellia luteilateralis (Covacevich & McDonald 1980) [=Concinnia luteilateralis, Tumbunascincus luteilateralis]C18
*Tachygia microlepis (Duméril & Bibron 1839)G74 [=Eumeces microlepisG74, *Liosoma microlepisM52, Lygosoma microlepisR13]
*Type species of generic name indicated
References
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