
Belongs within: Pteromalidae.
The Otitesellini are a group of wasps that oviposit on figs, either by entering the ostiole or through the fig wall. The larvae feed on the ovarium tissue, which swells to a gall (Bouček 1988).
The Sycoryctinae are a group of fig-wasps in which the females have a long, extruded ovipositor (at least partially covered by the extended last gastral tergite[s] in some Sycoryctini) and an undivided scutellum (Bouček 1988).
<==Otitesellini [Otitesellinae, Sycoecinae, Sycoecini, Sycoryctinae, Sycoryctini]BM22 |--PhilosycusB88 |--Eujacobsonia Grandi 1923B88 | `--*E. mirabilisB88 |--Guadalia Wiebes 1867B88 | `--*G. vissali Wiebes 1967B88 |--Micrognathophora Grandi 1921B88 | `--*M. leptopteraB88 |--Lipothymus Grandi 1921B88 | `--*L. sumatranusB88 |--Grandiana Wiebes 1961B88 | |--*G. wassae Wiebes 1867B88 | `--G. armadillo Bouček 1988B88 |--Micranisa Walker 1875 [incl. Epicolystichus Girault 1915, Sycobiella Westwood 1883]B88 | |--*M. pteromaloides [=Idarnes pteromaloides]B88 | |--M. aereicorpus (Girault 1915) [=*Epicolystichus aereicorpus]B88 | `--‘*Sycobiella’ saundersiiB88 |--Otitesella Westwood 1883MH11, B88 [incl. Paracolystichus Girault 1915B88] | |--*O. digitataB88 | |--O. amplipennis (Girault 1915) [=Paracolystichus amplipennis]B88 | |--O. claraeB88 | `--O. compressiventris (Girault 1915) [=*Paracolystichus compressiventris]B88 |--Walkerella Westwood 1883LXH13, B88 [incl. Asemantoideus Girault 1916B88, Terastiozoon Grandi 1921B88] | |--*W. temeraria [incl. Terastiozoon benjamini Joseph 1957]B88 | |--W. dubia [=*Asemantoideus dubius]B88 | |--‘*Terastiozoon’ jacobsoniB88 | |--W. kurandensis Bouček 1988B88 | `--W. yashiroi [=Otitesella yashiroi]B88 |--Apocrypta Coquerel 1855 (see below for synonymy)B88 | |--*A. perplexaB88 | |--A. acaeta Ulenberg in Ulenberg & Pelt 1985B88 | |--A. caudata (Girault 1915) [=*Goniogastrella caudatus]B88 | |--A. centrocampta Ulenberg in Ulenberg & Pelt 1985B88 | |--A. channingi (Girault 1913) [=*Idarnoides channingi]B88 | |--A. macroseta Ulenberg in Ulenberg & Pelt 1985B88 | |--A. mega Ulenberg in Ulenberg & Pelt 1985B88 | |--A. megischia Ulenberg in Ulenberg & Pelt 1985B88 | `--A. meromassa Ulenberg in Ulenberg & Pelt 1985B88 |--Dobunabaa Bouček 1988B88 | `--*D. flava Bouček 1988B88 |--Tenka Bouček 1988B88 | `--*T. percaudata Bouček 1988B88 |--Watshamiella Wiebes in Bouček, Watsham & Wiebes 1981 [=Metidarnes Riek 1970 (n. n.)]B88 | |--*W. alataB88 | `--W. aurea (Girault 1919) [=Philotrypesis aurea]B88 |--Philotrypesis Förster 1878 (see below for synonymy)B88 | |--*P. longicauda Förster 1878B88 | |--‘*Sycoscaptella’ affinisB88 | |--P. angela Girault 1915GM79 | |--‘Sycoscaptella’ anguliceps [=*Sycoscapterella anguliceps]B88 | |--P. aterrima (Saunders 1883) [=Idarnella aterrima]B88 | |--P. caricae [=Cynips caricae, *Idarnodes caricae]B88 | |--P. immaculata Girault 1915 [incl. P. longicauda Girault & Dodd in Girault 1915 non Förster 1878]B88 | |--‘*Philotrypomorpha’ indicaB88 | |--P. longiventris Girault 1915B88 | |--‘*Polanisa’ luteaB88 | |--‘Sycoscaptella’ quadrisetosa [=*Tetranemopteryx quadrisetosa]B88 | |--P. silvensis Girault 1915B88 | `--‘Idarnes’ transiens [=*Idarnella transiens]B88 |--Sycoscapter Saunders in Westwood 1883 (see below for synonymy)B88 | |--*S. insignisB88 | |--S. australis (Froggatt 1900)B88 [=Idarnis (l. c.) australisB88, Idarnes australisG28] | |--S. conocephalus Wiebes 1964B88 | |--S. huberi (Girault 1919) [=*Philotrypesella huberi, Sycoryctes huberi]B88 | |--‘*Sycorycteridea’ keralensisB88 | |--S. miltoni (Girault 1931) [=*Ideoidarnes miltoni]B88 | |--‘Sycoscapter’ monilifer [=*Sycoscapteridea monilifer]B88 | |--‘*Sycoryctes’ patellarisB88 | |--‘*Arachonia’ plumosaB88 | |--S. subaeneus (Girault 1915) [=*Idarnomorpha subaenea, Idarnes subaeneus]B88 | `--S. varicilia (Girault 1928) [=Sycoryctes varicilia]B88 |--CrossogasterB88 |--Sycoecus Waterston 1914BM22 |--Seres wardiBM22 |--Diaziella Grandi 1928B88 | |--*D. bicolorB88 | `--D. falcataB88 |--Robertsia Bouček 1988B88 | |--*R. mandibularis Bouček 1988B88 | `--R. xylosyciae Bouček 1988B88 |--MarginaliaBM22 |--AdiyodiellaBM22 |--BoucekaBM22 |--ComptoniellaBM22 |--Grasseiana Abdurahiman & Joseph 1967BM22, B88 | |--*G. callosaB88 | `--G. boschmai (Wiebes 1964) [=Sycobiella boschmai]B88 |--PhilosycellaBM22 `--PhiloverdanceBM22
Apocrypta Coquerel 1855 [incl. Agaonella Baker 1913, Goniogaster Mayr 1885, Goniogastrella Girault & Dodd in Girault 1915, Idarnoides Girault 1913; Apocryptini]B88
Philotrypesis Förster 1878 [incl. Idarnella Westwood 1883, Idarnodes Westwood 1883, Philotrypomorpha Abdurahiman & Joseph 1976, Polanisa Walker 1875 (nom. rej.), Sycoscaptella Westwood 1883, Sycoscapterella Ashmead 1904, Tetranemopteryx Ashmead 1904; Philotrypesini]B88
Sycoscapter Saunders in Westwood 1883 [incl. Arachonia Joseph 1957, Idarnomorpha Girault 1915, Ideoidarnes Girault 1931, Philotrypesella Girault 1919, Sycorycteridea Abdurahiman & Joseph 1975, Sycoryctes Mayr 1885, Sycoscapteridea Ashmead 1904]B88
*Type species of generic name indicated
References
[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).
[BM22] Burks, R., M.-D. Mitroiu, L. Fusu, J. M. Heraty, P. Janšta, S. Heydon, N. D.-S. Papilloud, R. S. Peters, E. V. Tselikh, J. B. Woolley, S. van Noort, H. Baur, A. Cruaud, C. Darling, M. Haas, P. Hanson, L. Krogmann & J.-Y. Rasplus. 2022. From hell’s heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera). Journal of Hymenoptera Research 94: 13–88.
[G28] Girault, A. A. 1928. A prodigeous discourse on wild animals. Privately published (reprinted: Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 218–220).
[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1–400.
[LXH13] Li, Z., H. Xiao & D.-W. Huang. 2013. Sirovena Bouček (Pteromalidae: Pireninae), a new member of the fig wasp community associated with Ficus microcarpa (Moraceae). Zootaxa 3619 (5): 581–588.
[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PloS One 6 (11): e27023.