Belongs within: Coreoidea.
Contains: Leptoscelis, Colpura, Leptoglossus, Coreus.
The Coreidae are a group of large plant-feeding bugs that typically produce strong repellent odours (Carver et al. 1991). The squash bug Anasa tristis is a pest of cucurbits. Species of Amblypelta and Theraptus cause premature fall of coconuts (Richards & Davies 1977).
Characters (from Carver et al. 1991): Stout, usually robust bugs with relatively thick legs; bucculae extending to, and usually behind, level of antennifers; head usually much less than half as wide as base of pronotum; ocelli present; metapleural scent gland openings with distinct peritremes; posterior margins of T4 and T5 produced posteriorly in mid line; membrane of fore wing with longitudinal accessory veins; corium without a large hyaline area.
<==Coreidae [Corizidae]
|--AgriopocorinaeCGW91
| |--AgriopocorisCGW91
| `--Tylocryptus egenusCGW91
|--CloresminaeD01a
| |--Notobitus abdominalis Distant 1901D01a
| `--CloresmusD01a
| |--C. khasianus Distant 1901D01a
| `--C. modestus Distant 1901D01a
|--PhysomerinaeD01b
| |--Rhyticoris spinipes [=Lygaeus spinipes; incl. Piezogaster patulus Walker 1871]D01b
| |--PhysomerusD01b
| | |--P. grossipes (Fabr. 1794) [=Lygaeus grossipes; incl. P. delineatus Walker 1871]D01b
| | `--P. subargenteus Walker 1871D01b
| `--AcanthocorisD01b
| |--A. anticus Walker 1871D01b
| |--A. esau Distant 1901D01b
| `--A. obscuricornis Dall. 1852 [incl. A. tarsalis Walker 1871]D01b
|--PseudophloeinaeCGW91
| |--GralliclavaCGW91
| |--ClavigralloidesCGW91
| |--Pseudophlaeus fallenii [=Coreus fallenii, Arenocoris fallenii]L49
| `--ClavigrallaD01b
| |--C. acantharis (Fabr. 1803) [=Lygaeus acantharis; incl. C. tuberculata Dall. 1852]D01b
| |--C. aliena (Walker 1871) [=Cletus alienus]D01b
| |--C. dispar Walker 1872 (n. d.)D01b
| |--C. gibbosaG01
| |--C. horrensD01b
| |--C. indecora (Walker 1871) [=Cletus indecorus]D01b
| `--C. spinigera Walker 1872 (n. d.)D01b
|--DiscogastrinaeD01b
| |--Savius Stål 1862 [incl. Homoeoceroides Walker 1871]D01b
| | `--‘Homoeocerus’ diversicornis [=*Homoeoceroides diversicornis]D01b
| |--CnemomisD01b
| | |--C. cognata Stål 1865 [incl. Hypselonotus signatus Walker 1871]D01b
| | `--C. dubia (Dall. 1852) [=Paryphes dubius; incl. Hypselonotus mundus Walker 1871]D01b
| |--ScamuriusD01b
| | |--S. amabilis (Stål 1855) [=Paryphes amabilis; incl. Homoeoceroides incongruus Walker 1871]D01b
| | `--S. scutellaris (Walker 1871) [=Hypselonotus scutellaris]D01b
| `--Discogaster Burmeister 1835 [incl. Leptornytus Walker 1871]D01b
| |--D. drewseniD01b
| `--D. rufocornis (Walker 1871) [=Leptornytus rufocornis]D01b
|--SpartocerinaeD01b
| |--SephinaD01b
| | |--S. humeralis Distant 1901D01b
| | |--S. limbata [incl. S. atra Walker 1871]D01b
| | `--S. vinula [incl. Spartocera lampyroides Walker 1871]D01b
| `--SpartoceraD01b
| |--S. batatas (Fabr. 1798) [=Lygaeus batatas]D01b
| |--S. brevicornisE66
| |--S. fusca (Thunb. 1783) [=Cimex fuscus; incl. Spartocerus diffusus]D01b
| |--S. giganteaD01b
| |--S. grandis Distant 1901D01b
| |--S. pantomima (Dist. 1881) [=Sephina pantomima; incl. Sp. ortonedai Montand. 1897]D01b
| `--S. rubicunda Spin. in Gay 1852 [incl. S. chilensis Walker 1871]D01b
|--GonocerinaeD01b
| |--GonocerusL-SSY04
| | |--G. annulipes Lucas 1847E12
| | |--G. insidiator [=Coreus insidiator]L49
| | `--G. juniperi Herrich-Schäffer 1839L-SSY04
| |--CletomorphaD01b
| | |--C. kirbyi Distant 1901 [=C. denticulata Kirby 1891 (preoc.)]D01b
| | |--C. raja Distant 1901D01b
| | `--C. walkeri Kirby 1891 [incl. C. benita Kirby 1891]D01b
| `--PlinacthusD01b
| |--P. acicularis (Fabr. 1803) [=Alydus acicularis; incl. Cletus conspicuus Walker 1871]D01b
| |--P. dubius (Herr.-Schäff. 1842) [=Gonocerus dubius; incl. Cletus clarus Walker 1871]D01b
| |--P. pungens (Thunb. 1783) [=Cimex pungens; incl. Gonocerus luridus Dall. 1852]D01b
| `--P. spinosus Stål 1859 [incl. Cletus mundus Walker 1871]D01b
|--AcanthocephalinaeD01a
| |--AcanthocephalaD01a
| | |--A. affinis (Walker 1871) [=Metapodius affinis]D01a
| | |--A. angustipes (Westw. in Hope 1842) [=Metapodius angustipes; incl. M. constrictus Walk. 1871]D01a
| | |--A. femorataL86
| | |--A. latipes (Drury 1782) [=Cimex latipes; incl. Metapodius sericeicollis Walk. 1871]D01a
| | `--A. unicolor (Westw. 1842) [=Metapodius unicolor; incl. M. distinctus Walk. 1871]D01a
| `--LaminicepsD01a
| |--L. cardinalisD01a
| |--L. ducalisD01a
| |--L. fasciatusD01a
| |--L. festivus Distant 1901D01a
| |--L. megaera (Burm. 1835) [=Paryphes megaera]D01a
| `--L. sumptuosus Distant 1901D01a
|--CentroscelinaeD01b
| |--Acidomeria strigata (Walker 1871) [=Gonocerus strigatus]D01b
| |--Namacus annulicornis Stål 1870 [incl. N. rufescens Walker 1872]D01b
| |--MargusD01b
| | |--M. inornatusD01b
| | `--M. obscurator (Fabr. 1803) [=Coreus obscurator]D01b
| |--CatorhinthaD01b
| | |--C. guttula (Fabr. 1794) [=Lygaeus guttula]D01b
| | `--C. semialba (Walker 1872) [=Namacus semialbus]D01b
| |--ParyphesD01b
| | |--P. festivus Costa 1863 [incl. P. gloriosus Walker 1871]D01b
| | `--P. viridipes Walker 1871 (n. d.)D01b
| `--HypselonotusD01b
| |--H. concinnus [incl. H. propinquus Walker 1871]D01b
| |--H. interruptus Hahn 1831 [incl. Jadera subvittata Walker 1871]D01b
| `--H. lanceolatus Walker 1871D01b
|--MictinaeD01b
| |--Saguntus lobulatus Stål 1865 [incl. Hypselonotus crassifemur Walker 1871]D01b
| |--Quintius pallens (Walker 1871) [=Hirileus pallens; incl. H. pallidus Walker 1871]D01b
| |--Mictis profana (Fabr. 1803)CGW91, D14 [=Lygaeus profanaD14; incl. M. profanus var. cruxK08]
| |--Molchina Amyot & Serv. 1843 [incl. Euplatycoris Walker 1871]D01b
| | `--M. granulata Stål 1870 [incl. Euplatycoris bellicornis Walker 1871]D01b
| |--ZorevaD01b
| | |--Z. marginalis (Walker 1871) [=Hypselonotus marginalis]D01b
| | `--Z. spiniferaD01b
| |--ArchimerusD01b
| | |--A. alternatusE66
| | |--A. calcaratorE66
| | `--A. pallens (Dall. 1852) [=Physomerus pallens, Piezogaster pallens]D01b
| `--NematopusD01b
| |--N. fasciatus Westw. in Hope 1842 [incl. Hypselonotus fascicollis Walker 1871]D01b
| |--N. indus (Linnaeus 1758)D01b [=Cimex (Oblongus) indusD01b, L58; incl. Hypselonotus pectoralis Walker 1871D01b]
| |--N. lepidus Stål 1862 [incl. Hypselonotus armatus Walker 1871]D01b
| `--N. ruficrus (Perty 1830) [=Anisoscelis ruficrus; incl. Hypselonotus ventralis Walker 1871]D01b
|--Leptoscelidini [Leptoscelinae]D01b
| |--LeptoscelisD01b
| |--Malvana minax (Walker 1871) [=Hypselonotus minax]D01b
| |--AmblyomiaB96a
| |--CoribergiaB96a
| |--DalmatomammuriusB96a
| |--KalinckascelisB96a
| |--MalvanaioidesB96a
| |--PlunentisB96a
| |--Sephinioides Brailovsky 1996B96a
| | `--*S. ugaldei Brailovsky 1996B96a
| `--PhthiaD01b
| |--P. affinis Distant 1901D01b
| |--P. cantharidinaD01b
| |--P. decorataD01b
| |--P. lunata [incl. P. concinna Walker 1871]D01b
| |--P. pictaD01b
| |--P. smaragdina (Walker 1871) [=Sphictyrtus smaragdinus]D01b
| `--P. ventralisD01b
|--ColpurinaeD01a
| | i. s.: SciophyrusD01a
| | |--S. anticus (Walker 1871) [=Cletus anticus]D01a
| | `--S. crassus (Walker 1871) [=Cletus crassus]D01a
| | PachycephalusD01a
| | |--P. opacusD01a
| | `--P. touchei Distant 1901D01a
| `--ColpuriniB96b
| |--ColpuraD01a
| |--AgathyrnaB96b
| |--Acanthotyla fasciata (Walker 1871)B96b, D01a [=Cletus fasciatusD01a]
| |--Monasavuhygia Brailovsky 1996B96b
| | `--*M. cordata Brailovsky 1996B96b
| |--Acarihygia Brailovsky 1993B96b
| | `--*A. fijiana Brailovsky 1993B96b
| `--Brachylybas Stål 1870B96b
| |--*B. variegata (Le Guillou 1841) [=Gonocerus variegatus, B. variegatus]B96b
| |--B. delgadoi Brailovksy 1996B96b
| |--B. taveuni Brailovsky 1996B96b
| `--B. vanua Brailovsky 1996B96b
|--CoreinaeCGW91
| |--Amblypelta cocophagaCGW91, MHG04
| |--Aulacosternum nigrorubrumCGW91, C91
| |--PompanotusCGW91
| |--TurranaCGW91
| |--LeptoglossusCGW91
| |--CoreusL02
| |--AmorbusCGW91
| | |--A. alternatusCGW91
| | `--A. rubiginosusG75
| `--CletusCGW91
| |--C. apicifer Walker 1871 (n. d.)D01b
| |--C. bipunctatus (Westw. in Hope 1842) [=Coreus bipunctatus; incl. Cl. signatus Walker 1871]D01b
| |--C. fuscescens Walker 1871D01b
| |--C. inconspicuus Walker 1871D01b
| |--C. punctiger (Dall. 1852) [=Gonocerus punctiger; incl. Homoeocerus minax Walker 1871]D01b
| |--C. punctulatus (Westw. in Hope 1842) [=Coreus punctulatus; incl. Cl. femoralis Kirby 1891]D01b
| |--C. rubidiventris (Wastw. in Hope 1842) [=Coreus rubidiventris; incl. Cl. pallescens Walker 1871]D01b
| |--C. similisWEE70
| `--C. subnotatus Walker 1871D01b
|--CorizinaeD01b
| |--CorizusH01
| | |--C. crassicornis [incl. C. crassicornis var. abutilon]H01
| | |--C. maculatusH01
| | |--C. parumpunctatusH01
| | `--C. subrufus [incl. C. capitatus]H01
| |--JaderaD01b
| | |--J. abdominalis Walker 1871 (n. d.)D01b
| | |--J. aeola (Dallas 1852) [=Serinetha aeola; incl. Lygaeus conspersus Walker 1872]D01b
| | |--J. antica (Walker 1872) [=Lygaeus anticus]D01b
| | |--J. haematoloma [incl. Lygaeus marginalis Walker 1872]D01c
| | `--J. sanguinolenta (Fabr. 1775) [=Cimex sanguinolentus; incl. Lygaeus rufoculis Kirby 1890]D01b
| `--SerinethaD01b
| |--S. abdominalis (Fabr. 1803) [=Lygaeus abdominalis; incl. S. taprobanensis Dallas 1852]D01b
| |--S. aethiops Dallas 1901D01b
| |--S. fraternaD01b
| |--S. lurida Dallas 1852D01b
| |--S. subrufescens (Kirby 1888)D01c [=Lygaeus subrufescensD01c, Leptocoris subrufescensJG19]
| | |--S. s. subrufescensJG19
| | `--‘Leptocoris’ s. flavusJG19
| `--S. vicina Dallas 1852 [incl. S. coxalis Kirby 1891, Astacops nigricornis Walker 1872]D01b
`--Homoeocerus [Homoeocerinae]D01a
|--H. abbreviatus (Fabr. 1794) [=Lygaeus abbreviatus; incl. H. umbratus Walker 1871]D01a
|--H. abdominalis Distant 1901D01a
|--H. angulatus West. in Hope 1842 (see below for synonymy)D01a
|--H. annulatus (Thunb. 1822) [=Alydus annulatus; incl. Ceratopachys subsparsus Walker 1871]D01a
|--H. atkinsoni Distant 1901D01a
|--H. auriculatusD01a
|--H. borneensis Distant 1901D01a
|--H. concisus Walker 1871D01a
|--H. graminis (Fabr. 1803) [=Lygaeus graminis; incl. H. turbidus Walker 1871]D01a
|--H. lineatus Walker 1871 (n. d.)D01a
|--H. lucidus Walker 1871D01a
|--H. magnicornisD01a
|--H. malayensis Distant 1901D01a
|--H. montanus Distant 1901D01a
|--H. nigridorsumD01a
|--H. prominulus (Dall 1852) [=Ceratopachys prominulus; incl. C. vicinus Sall 1852]D01a
|--H. serrifer (Westw. in Hope 1842) (see below for synonymy)D01a
|--H. signatus Walker 1871 [incl. H. biplagiatus Stål 1873]D01a
|--H. simplex Walker 1871D01a
|--H. sinicus Walker 1871D01a
|--H. smecticus Distant 1901D01a
|--H. subjectus Walker 1871D01a
|--H. tinctusD01a
|--H. trimaculatus Distant 1901D01a
|--H. unipunctatus (Thunb. 1783) [=Cimex unipunctatus; incl. H. chinensis Dall 1852]D01a
|--H. variabilis (Dall 1852) [=Ceratopachys variabilis; incl. H. anticus Walker 1871]D01a
|--H. walkerianus [incl. H. plagiatus Walker 1871 (preoc.)]D01a
`--H. yerburyi Distant 1901D01a
Coreidae incertae sedis:
Paradasynus spinosusYS01
Thasus gigasGE05, EN20
Narnia inornataI92
Aufeius impressicollisBM76
SyromastesH01
|--S. marginatusH01
`--S. spinigera [=Coreus spiniger]L49
Therapha hyoscyami (Linnaeus 1758)H01, L58 [=Cimex (Oblongus) hyoscyamiL49, L58, Corizus hyosciamiL49]
EnoplopsH01
|--E. scaphaH01 [=Coreus scaphaL49, Syromastes scaphaL49]
`--E. sibiricaH01
ChelonideaG75
|--C. tabulataG75
`--C. vittigerG75
Anasa tristisCGW91, RD77
HebeicorisYR12
|--H. longaYR12
|--H. luanpingensisYR12
`--H. xinboensis Hong 1985RJ93
Uranocoris Walker 1871D01b
`--U. suavis Walker 1871D01b
Menenotus diminutus Walker 1871D01b
Derepteryx truncata Walker 1871D01b
Hyalymenus walkeri [incl. H. ichneumoniformis Walker 1871 (preoc.)]D01b
VerlusiaL49
|--V. quadrata [=Coreus quadratus, Syromastes quadratus]L49
|--V. rhombeaD01b [=Coreus rhombeusL49, Cimex rhombeusL49]
`--V. sulcicornis [=Coreus sulcicornis; incl. V. rotundiventris]L49
Dulichius inflatusRD77
TheraptusRD77
Chariestrus antennatorE66
Pachylis Lepeletier & Serville 1825E66, K18
`--P. argentinusE66
SinocorisYR12
|--S. oblongaYR12
`--S. ovataYR12
Karatavocoris asiaticaYR12
Corioides fortisYR12
Kezuocoris liaoningensisYR12
Kerjiecoris oopsisYR12
Weichangicoris daobaliangensisYR12
Bibiticen hebeiensisYR12
Phyllomorpha algirica Guér. 1839 [incl. P. laciniata]L49
Micrelytra fossularum (see below for synonymy)L49
Astemma aptera (Linnaeus 1758)L49, L58 (see below for synonymy)
Astemma aptera (Linnaeus 1758)L49, L58 [=Cicada (Spumans) apteraL49, L58, Eurycephala apteraL49; incl. Salda pallicornisL49, Acanthia pallicornisL49, Halticus pallicornisL49, Phytocoris pallicornisL49]
Homoeocerus angulatus West. in Hope 1842 [incl. H. clarus Walker 1871, H. mundus Walker 1871, H. parallelus Walker 1871]D01a
Homoeocerus serrifer (Westw. in Hope 1842) [=Coreus serrifer; incl. H. parvulus Walker 1871, H. unipunctatus Dall 1852 non Cimex unipunctatus Thunb. 1783]D01a
Micrelytra fossularum [=Hydrometra fossularum, Arctorus fossularum, Micrelytrum fossularum, Stenocephalus fossularum; incl. Alydus apterus]L49
*Type species of generic name indicated
References
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[B96a] Brailovsky, H. 1996a. A new genus and a new species of Leptoscelidini (Heteroptera: Coreidae) from Costa Rica. Proceedings of the Entomological Society of Washington 98 (2): 215–220.
[B96b] Brailovsky, H. 1996b. A revision of the tribe Colpurini (Heteroptera: Coreidae) from the Fiji islands. Proceedings of the Entomological Society of Washington 98 (3): 473–490.
[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).
[C91] CSIRO. 1991. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1. Melbourne University Press: Carlton (Victoria).
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[YS01] Yoshizawa, K., & T. Saigusa. 2001. Phylogenetic analysis of paraneopteran orders (Insecta: Neoptera) based on forewing base structure with comments on monophyly of Auchenorrhycha (Hemiptera). Systematic Entomology 26: 1–13.