Costispiriferidae

Belongs within: Spiriferida.

The Costispiriferidae are a Devonian family of brachiopods with a biconvex shell in which a fold and sulcus is not well developed (Boucot et al. 1965).

Characters (Boucot et al. 1965): Biconvex, weakly to moderately transverse; lateral slopes with moderately to very numerous simple costae; fold and sulcus generally indistinct, poorly delineated from lateral slopes, provided with costae which are indistinguishable from lateral costae except for tendency toward bifurcation; microornament of distinct capillae and weak growth lamellae in some; pedicle valve interior with dental plates, lacking delthyrial plate and median septum; brachial valve interior with longitudinally striated area of diductor attachment, with or without crural plates; shell substance impunctate.

<==Costispiriferidae [Costispiriferinae, Theodossiinae, Theodossioidea]BJ65
    |--Lazutkinia Rzhonsnitskaya 1951 [incl. Yavorskiella Lazutkin in Yavorsky 1940 (n. n.)]BJ65
    |    `--*L. mamontoviensis (Lazutkin in Yavorsky 1940) [=Spirifer (Yavorskiella) mamontoviensis]BJ65
    |--Urella Rzhonsnitskaya 1960BJ65
    |    `--*U. asiatica Rzhonsnitskaya 1960BJ65
    |--Eudoxina Frederiks 1929BJ65
    |    |--*E. media (Lebedev 1912) [=Spirifer medius]BJ65
    |    `--E. subrotundaHB93
    |--Costispirifer Cooper 1942BJ65
    |    |--*C. planicostatus (Swartz 1929) [=Spirifer arenosus planicostatus]BJ65
    |    `--C. arenosus (Conrad 1841)HB93
    `--Theodossia Nalivkin 1925 [incl. Vandergrachtella Crickmay 1953]BJ65
         |--T. anossofi (de Verneuil 1845) [=Spirifer anossofi]BJ65
         |--T. arcuum (Crickmay 1953) [=*Vandergrachtella arcuum]BJ65
         |--T. hungerfordiBJ65
         `--T. keeneiN79

*Type species of generic name indicated

References

[BJ65] Boucot, A. J., J. G. Johnson, C. W. Pitrat & R. D. Staton. 1965. Spiriferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt H. Brachiopoda vol. 2 pp. H632–H728. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas Press: Lawrence (Kansas).

[HB93] Harper, D. A. T., C. H. C. Brunton, L. R. M. Cocks, P. Copper, E. N. Doyle, A. L. Jeffrey, E. F. Owen, M. A. Parkes, L. E. Popov & C. D. Prosser. 1993. Brachiopoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 427–462. Chapman & Hall: London.

[N79] Norris, A. W. 1979. Devonian in the Western Hemisphere. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A218–A253. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).