Eurytomidae

 Buresium naso, photographed by Mike Gates.

Belongs within: Chalcidoidea.
Contains: Eurytominae.

The Eurytomidae are a family of chalcidoid wasps including both parasitic and phytophagous species. The pronotum in eurytomids is large and subrectangular, and the majority of species are non-metallic in coloration and bear strong piliferous puncturation (Bouček 1988).

Characters (from Bouček 1988): Body usually non-metallic, black or marked with yellow or reddish; thorax mostly with coarse piliferous puncturation; pronotum subrectangular, large; notauli complete though posteriorly sometimes shallow; lower mesepimeron not separated; forewing without linea calva; tarsi five-segmented; gaster convex, non-collapsing, ovipositor hardly exserted (if apex narrowed in a cauda then covered by extended epipygium).

<==EurytomidaeHB13
    |--BuresiinaeHB13
    |    |--MacrorileyaHB13
    |    `--BuresiumHB13
    |         |--B. nasoHB13
    |         `--B. rufumHB13
    `--+--EurytominaeHB13
       `--Rileyinae [Rileyini]B88
            |  i. s.: ArchirileyaB88
            |         Heimbrella Subba Rao 1978B88
            |         Calorileya Crawford 1910B88
            |         Dougiola Bouček 1988B88
            |           `--*D. koebelei (Ashmead 1900) [=Systole koebelei]B88
            |         Matna Bouček 1988B88
            |           `--*M. fallax Bouček 1988B88
            |--+--Neorileya flavipesHB13
            |  `--Heimbra [Heimbrinae, Heimbrini]HB13
            |       `--H. opacaHB13
            `--Rileya Ashmead 1888 (see below for synonymy)B88
                 |  i. s.: *R. cecidomyiaeB88
                 |         R. asiaticaB88
                 |         R. spadix (Girault 1915) [=*Pararileya spadix]B88
                 |         R. vardyiB88
                 |--+--R. atopogasterMH11
                 |  `--R. grisselliMH11
                 `--+--R. longitergumHB13
                    `--+--R. pallidipesHB13
                       `--Austrophotismus Girault 1938HB13, B88
                            `--A. daicles (Walker 1839) [=Miscogaster daicles; incl. *A. ater Girault 1938]B88
Eurytomidae incertae sedis:
  Sinoeurytomites Hong 2002H02
    `--*S. nigratus Hong 2002H02
  Eoeurytomites Hong 2002H02
    `--*E. badius Hong 2002H02
  Zhuia Hong 2002H02
    `--*Z. minor Hong 2002H02
  Liaoeurytomites Hong 2002H02
    `--*L. petiolatus Hong 2002H02
  Fushuneurytomites Hong 2002H02
    `--*F. xilutianensis Hong 2002H02
  BephratelloidesGF96
    |--B. ablusus Grissell & Foster 1996GF96
    |--B. cubensisGF96
    |--B. limai (n. d.)GF96
    |--B. melleus (n. d.)GF96
    |--B. paraguayensisGF96
    |--B. petiolatusGF96
    `--B. pomorumGF96
  ChryseidaB88

Rileya Ashmead 1888 [incl. Ashmeadia Howard 1889, Pararileya Girault 1915, Tragiicola Brethes 1918, Xenopelte Cameron 1913]B88

*Type species of generic name indicated

References

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[GF96] Grissell, E. E., & M. S. Foster. 1996. A new Bephratelloides (Hymenoptera: Eurytomidae) from seeds of Cymbopetalum (Annonaceae) in Mexico. Proceedings of the Entomological Society of Washington 98 (2): 256–263.

[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466–542.

[H02] Hong Y. 2002. Amber Insect of China. Beijing Scientific and Technological Publishing House.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PloS One 6 (11): e27023.

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