Belongs within: Meliphagoidea.
Contains: Gerygone, Acanthiza.
The Acanthizidae, Australo-Papuan warblers, are a group of small, generally dull-coloured birds. Most species have short rounded wings and a fairly short tail; terrestrial species have relatively strong legs and feet (Internet Bird Collection). The goldenface Pachycare flavogriseum of New Guinea is more boldly coloured than other species, with a yellow face and underparts and grey upperparts.
Characters (Norman et al. 2009): Skull with dorsal-most point of profile roughly level with caudal border of orbit; lateral profile of top of skull with strongly marked slope from apex to nasofrontal hinge, profile thus lacking horizontal component to the dorsal surface; humerus not extending far distally relative to position of fossa pneumotricipitalis, crista deltopectoralis deflected cranially ; tarsometatarsus with trochleae II and IV both ≤50% of width of trochlea III, trochlea II projecting outwards slightly at base then straightening over distal length to be parallel with medial border, trochlea IV straight and confluent with lateral border, incisurae trochlearis lateralis and medialis both of about equal depth.
<==Acanthizidae [Acanthizeae, Acanthizinae]
|--+--Oreoscopus gutturalisBKB15
| `--Pachycare flavogriseumBKB15
`--+--GerygoneBKB15
`--+--+--AcanthizaBKB15
| `--+--Acanthornis magnaBKB15
| `--Aphelocephala Oberholser 1899BKB15, B94 [Aphelocephalinae]
| |--A. leucopsis (Gould 1840) [=Xerophila leucopsis]WS48
| | |--A. l. leucopsisWS48 [incl. A. l. whiteiM03]
| | `--A. l. castaneiventris (Milligan 1903) (see below for synonymy)WS48
| |--A. nigricincta (North 1895) [=Xerophila nigricincta]WS48
| `--A. pectoralisJT12
`--+--+--+--Pycnoptilus floccosusBKB15
| | `--+--ChthonicolaJT12
| | | |--C. minimaN87
| | | `--C. sagittatusJT12
| | `--PyrrholaemusBKB15
| | |--P. brunneus Gould 1840 (see below for synonymy)WS48
| | `--P. sagittatusBKB15
| `--+--Hylacola Gould 1842BKB15, G42
| | |--*H. pyrrhopygia (Vigors & Horsfield 1827)G42, WS48 [=Acanthiza pyrrhopygiaWS48]
| | `--H. cauta Gould 1842BKB15, WS48
| | |--H. c. cautaM03 (see below for synonymy)
| | |--H. c. halmaturinusM03
| | `--H. c. macrorhynchusM03
| `--CalamanthusBKB15
| |--C. campestris (Gould 1840)WS48 [=Praticola campestrisWS48, C. fuliginosus campestrisS90]
| | |--C. c. campestrisWS48
| | `--C. c. isabellinus North 1896 (see below for synonymy)WS48
| |--C. cautusAP21
| |--C. fuliginosus (Vigors & Horsfield 1827) [=Anthus fuliginosus]WS48
| |--C. montanellus Milligan 1903AP21, WS48 (see below for synonymy)
| `--C. pyrrhopygiusAP21
`--+--Smicrornis Gould 1842BKB15, G42
| `--S. brevirostris (Gould 1838) [=Psilopus brevirostris]WS48
| |--S. b. brevirostrisWS48
| |--*S. b. flavescens Gould 1842G42, WS48 (see below for synonymy)
| |--S. b. occidentalis Sharpe 1879 [incl. S. brevirostris stirlingi Mathews 1912]WS48
| `--S. b. ochrogasterM03
`--+--+--OrigmaBKB15
| | |--O. rubricataN87
| | `--O. solitariaBKB15
| `--CrateroscelisBKB15
| |--C. murinaJT12
| |--C. nigrorufaJT12
| `--C. robustaBKB15
`--Sericornis Gould 1838BKB15, B94 [Sericornithinae]
| i. s.: S. arfakianusJT12
| S. beccariiJT12
| |--S. b. beccarii [=S. magnirostris beccarii]M03
| |--S. b. dubius [=S. magnirostris dubius]M03
| `--S. b. minimusM03
| S. cautusSJ85
| S. citreogularisAP21
| |--S. c. citreogularisM03
| |--S. c. cairnsiM03
| `--S. c. intermediusM03
| S. fuliginosusSJ85
| S. maculatus Gould 1847WS48 (see below for synonymy)
| S. magnusG68
| S. rufescensJT12
| S. spiloderaJT12
| S. virgatusJT12
|--+--S. papuensisBKB15
| `--S. perspicillatusBKB15
`--+--+--S. magnirostraBKB15
| | |--S. m. magnirostraM03
| | |--S. m. howeiM03
| | `--S. m. viridiorM03
| `--S. nouhuysiBKB15
`--+--S. frontalisBKB15
| |--S. f. frontalisM03
| |--S. f. ashbyiM03
| |--S. f. flindersiM03
| |--S. f. laevigasterM03
| |--S. f. melloriM03
| |--S. f. rosinaeM03
| |--S. f. tregellasiM03
| `--S. f. tweediM03
`--+--S. humilisBKB15 [=S. frontalis humilisM03]
`--S. keriBKB15
Aphelocephala leucopsis castaneiventris (Milligan 1903) [=Xerophila castaneiventris; incl. A. castaneiventris minilya Mathews 1920]WS48
Calamanthus campestris isabellinus North 1896 [incl. C. campestris peroni Mathews 1917, C. rubiginosus Campbell 1899, C. campestris wayensis Mathews 1912]WS48
Calamanthus montanellus Milligan 1903AP21, WS48 [=Calamanthus fuliginosus montanellusWS48; incl. C. montanellus ashbyi Mathews 1922WS48, C. fuliginosus carteri Mathews 1912WS48, C. campestris dorrie Mathews 1912WS48, C. campestris hartogi Carter 1916WS48, C. montanellus leakei Mathews 1922WS48]
Hylacola cauta cauta Gould 1842WS48 [=H. pyrrhopygia cautaWS48; incl. H. pyrrhopygia whitlocki Mathews 1912WS48, H. cauta whitlockiM03]
Pyrrholaemus brunneus Gould 1840 [incl. P. brunneus kalgoorlie Mathews 1922, P. brunneus milligani Mathews 1922, Sericornis brunnea pallescens Mathews 1912]WS48
Sericornis maculatus Gould 1847WS48 [=S. frontalis maculatusM03; incl. S. balstoni Ogilvie-Grant 1909WS48, S. frontalis balstoniM03, S. maculata condoni Mathews 1942WS48, S. maculatus fuscipes Alexander 1922WS48, S. maculatus geraldtonensis Mellor 1921WS48, S. maculatus hartogi Carter 1916WS48, S. maculatus houtmanensis Zietz 1921WS48, S. maculata mondraini Mathews 1942WS48, S. maculatus warreni Mathews 1912WS48]
*Smicrornis brevirostris flavescens Gould 1842G42, WS48 [incl. S. brevirostris mungi Mathews 1912WS48, S. brevirostris rogersi Mathews 1912WS48]
*Type species of generic name indicated
References
[AP21] Arlott, N., & B. van Perlo. 2021. Collins Birds of the World. William Collins.
[B94] Bock, W. J. 1994. History and nomenclature of avian family-group names. Bulletin of the American Museum of Natural History 222: 1–281.
[BKB15] Burleigh, J. G., R. T. Kimball & E. L. Braun. 2015. Building the avian tree of life using a large-scale, sparse supermatrix. Molecular Phylogenetics and Evolution 84: 53–63.
[G42] Gould, J. 1842. Descriptions of thirty new species of birds from Australia. Proceedings of the Zoological Society of London 10: 131–140.
[G68] Grant, P. R. 1968. Bill size, body size, and the ecological adaptations of bird species to competitive situations on islands. Systematic Zoology 17 (3): 319–333.
[JT12] Jetz, W., G. H. Thomas, J. B. Joy, K. Hartmann & A. Ø. Mooers. 2012. The global diversity of birds in space and time. Nature 491: 444–448.
[M03] Morcombe, M. 2003. Field Guide to Australian Birds 2nd ed. Steve Parish Publishing.
[N87] North, A. J. 1887. List of references to authentic descriptions of Australian birds’ eggs. Proceedings of the Linnean Society of New South Wales, series 2, 1 (4): 1163–1174.
[S90] Storr, G. M. 1990. Birds of Shark Bay area, Western Australia. In: Berry, P. F., S. D. Bradshaw & B. R. Wilson (eds) Research in Shark Bay: Report of the France-Australe Bicentenary Expedition Committee pp. 299–311. Western Australian Museum.
[SJ85] Storr, G. M., & R. E. Johnstone. 1985. Field Guide to the Birds of Western Australia 2nd ed. Western Australian Museum: Perth (Australia).
[WS48] Whittell, H. M. & D. L. Serventy. 1948. A systematic list of the birds of Western Australia. Public Library, Museum and Art Gallery of Western Australia, Special Publication 1: 1–126.