Belongs within: Acteonacea.
Contains: Pupa.
The Acteonidae are a group of marine gastropods of uncertain affinities. Though anatomically similar to opisthobranchs, they possess certain plesiomorphic features such as the presence of an operculum to close the shell aperture. The shell is well calcified with a distinct spire and an elongate aperture. The radula usually lacks a rachidian tooth and possesses from five to over two hundred laterals each side. Members of the subfamily Liocareninae differ from the Acteoninae in the absence of punctate sculpture and the presence of varices.
Acteonidae [Pupacea, Pupidae, Tornatelladae, Tornatellidae, Tornatellinae]
|--Liocarenus Harris & Burrows 1891 [Liocareninae]BR05
| `--*L. conovuliformis (Deshayes 1824) [=Auricula conovuliformis]BR17
`--Acteoninae [Actaeoninae]BR05
| i. s.: Eoactaeon Stephenson 1955KB01
| Nucleopsis Conrad 1865 [Nucleopsinae]BR05
| `--*N. subvaricata (Conrad 1860) [=Acteonina subvaricata]BR17
| Tornatellaea Conrad 1860 [Tornatellaeinae]BR05
| |--*T. bella Conrad 1860BR17
| |--T. cretacea Wade 1926KB01
| |--T. fontis (Duméril 1806)TTE93
| |--T. globulosa Wade 1926KB01
| `--T. hebertiTTE93
| Tornatella Lamarck 1816BR05
| |--*T. flammea (Bruguière 1789)BR17 (see below for synonymy)
| |--T. bullataR42
| |--T. glabra Reeve 1842R42
| |--T. minuta Petterd 1879P79
| |--T. oryza Reeve 1842R42
| |--T. ovalisH86
| |--T. siebaldii Reeve 1842R42
| |--T. tessellata Reeve 1842R42
| |--T. venustaR42
| `--T. virgata Reeve 1842R42
| Acteon Montfort 1810 [=Actaeon (l. c.) non Rang 1829]BR05
| |--*A. tornatilis (Linnaeus 1758)BR17 [=Bulla tornatilisBR17, Bulimus tornatilisR42, Voluta tornatilisKBP02]
| |--A. affinis (Sowerby 1836) [=Tornatella affinis]F26
| |--A. ambiguus (Hutton 1885) [=Admete ambigua]PH90
| |--‘Actaeon’ articulatusD56
| |--A. chattonensisPB29
| |--A. cicatricosus Sohl 1964KBP02
| |--A. cratericulatus Hedley 1906P61 [=A. craticulatus Murdoch & Suter 1906F26]
| |--A. fasciatusO27 [=Tornatella fasciataO27; incl. Auricula bifasciataR42, T. bifasciataG20]
| |--‘Actaeon’ muelleri Bosquet 1861KB01
| |--A. oneroaensis Powell & Bartrum 1929PB29
| |--A. ooliticusKBP02
| |--A. panamensis Dall 1908PP78
| |--A. praestitus Finlay 1924 [=A. sulcatus (Hutton 1885) (preoc.)]F27
| |--A. roseus Hedley 1906F26
| |--A. tenellusN79
| `--A. traski Stearns 1898O27
|--PupaBSK21
`--+--Rictaxis Dall 1871BSK21, O27
| |--R. painei (Dall 1903) [=Acteon (Rictaxis) painei]O27
| `--R. punctocoelatus (Carpenter 1864)BSK21, O27 [=Acteon (Rictaxis) punctocoelataO27, Tornatella punctocoelataO27]
| |--R. p. punctocoelatusO27
| `--R. p. vancouverensis (Oldroyd 1927) [=Acteon (Rictaxis) punctocoelata vancouverensis]O27
`--Aplustridae [Bullinadae, Bullinidae]BR05
|--Palaeohydatina flouestiTTE93
|--Hydatina Schumacher 1817BSK21, BR05
| `--H. physis (Linnaeus 1758)P61 [=Bulla physisP61, Bullina physisBR17, *Hydatina filosa Schumacher 1817BR17]
|--Nonacteonina Stephenson 1941 [Nonacteoninidae]BR05
| |--*N. graphoides Stephenson 1941BR17 [=Crenilabium (*Nonacteonina) graphoidesBS00]
| |--N. chilensis (Orbigny 1846) [=Scaphander chilensis, Crenilabium (Nonacteonina) chilensis]BS00
| `--N. deflexa (Stephenson 1941) [=Crenilabium (Nonacteonina) deflexa]BS00
|--Aplustrum Schumacher 1817 [=Amplustrum (l. c.); Amplustrina, Aplustrinae]BR05
| `--*A. fasciatum Schumacher 1817BR17
|--Sulcoactaeon Cossmann 1895 [Sulcoactaeonidae]BR05
| |--*S. striatosulcata (Zittel & Goubert 1861) [=Acteonina striatosulcata]BR17
| `--S. pulloides (Hudleston 1896)GK02
`--Bullina Férussac 1822BR17, BR05
|--B. amplustre (Linnaeus 1758)BR17, L58 [=Bulla amplustreBR17, Aplustrum aplustreH09]
|--B. lauta Pease 1860K65
|--B. undulata [=Bulla undulata]BR17
|--B. velum [=Bulla velum]BR17
`--B. vitrea Pease 1860 [=Bulla vitrea non Adams 1850 (not preoc. if in dif. gen.)]K65
Acteonidae incertae sedis:
Dactylus Schumacher 1817BR05
|--D. maurusF66
`--D. mustelinusF66
Microglyphis Dall 1902O27
|--*M. curtulus [=Acteon curtulus]O27
|--M. breviculus Dall 1902O27
|--M. estuarinus Dall 1908O27
`--M. globularis (Smith 1875) [=Admete globularis]PH90
Bullinula Swainson 1840P61
`--*B. lineata (Gray 1825)P61 (see below for synonymy)
Neactaeonina Thiele 1912P61
|--*N. cingulata [=Acteonina cingulata]P61
`--N. inexpectata Dell 1956P61
Crenilabium Cossmann 1889BS00
`--C. (Eoacteon Stephenson 1955)BS00
|--C. (*E.) linteus (Conrad 1858) [=Solidulus linteus, Actaeon linteus]BS00
`--C. (E.) valdovinosensis Bandel & Stinnesbeck 2000BS00
Japonacteon suturalis (Adams 1855)WBS-S15
Lissactaeon exilisP71
*Bullinula lineata (Gray 1825)P61 [=Bulla lineataP61, Bullina lineataBR17; incl. Bulla scabra Gmelin 1791 (preoc.)BR17, F26, Bullina scabraF26, Voluta ziczac Muhlfeldt 1878 non Schroeter 1804F26]
*Tornatella flammea (Bruguière 1789)BR17 [=Buccinum flammeumBR17, Actaeon flammeusH09, Voluta flammeaR42; incl. Bulimus variegatusR42]
*Type species of generic name indicated
References
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