Alveolinidae

Cross-section of Fasciolites boscii, from here.

Belongs within: Miliolina.

The Alveolinidae are a group of porcelaneous, often large (up to 100 mm in length) Foraminifera known from the Lower Cretaceous to the present (Loeblich & Tappan 1964). However, a significant gap exists in the fossil record between Praealveolina and other Cretaceous genera, and the Cenozoic members of the family (BouDagher-Fadel 2008).

Characters (BouDagher-Fadel 2008): Test: free, large, planispiral to fusiform, subcylindrical or globular, coiled about elongate axis. Early whorls may be irregular, streptospiral in monomorphic species, may be restricted to microspheric forms in dimorphic species. Chamber subdivided into chamberlets by longitudinal partitions (septula) perpendicular to main septa, and connected by passages below apertural face. Basal layer may include anastomosing canals. Aperture a slit parallel to base of apertural face, or single row of circular openings, or numerous rows of such openings in horizontal rows, arranged in definite patterns matching patterns of chamber divisions.

Alveolinidae (see below for synonymy)
|--Ovalveolina Reichel 1936LT64
| `--*O. ovum (d’Orbigny 1850) [=Alveolina ovum]LT64
|--Subalveolina Reichel 1936LT64
| `--*S. dordonica Reichel 1936LT64
|--Bullalveolina Reichel 1936LT64
| `--*B. bulloides (d’Orbigny in de la Sagra 1839) [=Alveolina bulloides]LT64
|--Cisalveolina Reichel 1941LT64
| `--*C. fallax Reichel 1941LT64
|--Multispirina Reichel 1947LT64
| `--*M. iranensis Reichel 1947LT64
|--Globoreticulina Rahaghi 1978B-F08
| `--G. iranicaB-F08
|--Praebullalveolina Sirel & Acar 1982B-F08
| `--P. afyonicaB-F08
|--Archaealveolina Fourcade 1980HW93
| `--A. reicheliHS02
|--Flosculinella Schubert in Richarz 1910LT64
| |--*F. bontangensis (Rutten 1913)LT64 [=Alveolinella bontangensisLT64, Flosculina bontangensisC40]
| `--F. reicheliM12
|--Alveolinella Douvillé 1906LT64
| |--*A. quoyi (d’Orbigny 1826) [=Alveolina quoii]LT64
| |--A. bosciiC40
| `--A. praequoyiM12
|--Glomalveolina Hottinger 1962B-F08
| |--*G. dachelensis (Schwager 1883) [=Alveolina dachelensis, Fasciolites (*Glomalveolina) dachelensis]LT64
| |--G. delicatissimaB-F08
| `--G. primaeva [=Fasciolites (Glomalveolina) primaeva]LT64
|--Borelis de Montfort 1808 [=Borelia Agassiz 1844]LT64
| |--*B. melonoides de Montfort 1808LT64
| |--B. haueriB-F08
| |--B. pygmaeusB-F08
| `--B. schlumbergeriPHT13
|--Praealveolina Reichel 1933LT64
| |--P. (Praealveolina)LT64
| | |--*P. (P.) tenuis Reichel 1933LT64
| | `--P. (P.) cretacea (d’Archiac 1837) [=Alveolina cretacea]LT64
| `--P. (Simplalveolina Reichel in Loeblich & Tappan 1964)LT64
| `--P. (*S.) simplex Reichel 1936LT64
|--Fasciolites Parkinson 1811LT64 (see below for synonymy)
| |--*F. oblonga (d’Orbigny 1826) [=Alveolina oblonga]LT64
| |--F. boscii (Defrance in Bronn 1825) [=*Oryzaria boscii]LT64
| |--‘Flosculina’ daunica Checchia-Rispoli 1912 [=F. (*Checchiaites) daunica]LT64
| |--*Flosculina’ decipiens Schwager 1883LT64
| |--‘Flosculina’ globosaC40 [=Alveolina globosaB-F08]
| |--F. levantinaLT64
| |--F. pasticillatusLT64
| |--*Miliolites’ sabulosus de Montfort 1808LT64
| |--F. schwageriLT64
| `--‘Alveolina’ violae Checchia-Rispoli 1912 [=*Eoalveolinella violae]LT64
`--Alveolina d’Orbigny 1826B-F08 (see below for synonymy)
|--*A. melo (Fichtel & Moll 1798) (see below for synonymy)LT64
|--A. aramaeaB-F08
|--A. elliptica [incl. A. elliptica var. flosculina]B-F08
| |--A. e. ellipticaB-F08
| `--A. e. nuttalliB-F08
|--A. elongataM12
|--A. fragilisM12
|--A. fusiformisM12
|--A. katicaeB-F08
|--A. minervensisB-F08
|--A. subpyrenaicaB-F08
`--A. vredenburgi Davies 1937 [incl. A. cucumiformis Hottinger 1960]MP14

Nomen nudum: Alveolina ovulum Stache in Schwager 1883 [=A. (Glomalveolina) ovulum]LT64

Alveolina d’Orbigny 1826B-F08 [incl. Clausulus de Montfort 1808LT64, Melonia Lamarck 1822LT64, Melonites Lamarck 1812LT64, Neoalveolina Silvestri 1928LT64]

*Alveolina melo (Fichtel & Moll 1798) [=Nautilus melo, Borelis melo; incl. A. bradyi Silvestri 1927, *Neoalveolina bradyi, *Clausulus indicator de Montfort 1808, *Melonites sphaerica Lamarck 1816, *Melonia sphaerica]LT64

Alveolinidae [Alveolinea, Alveolinellidae, Alveolinellinae, Alveolinida, Alveolininae, Borelida, Borelidae, Borelidinae, Fasciolitidae]

Fasciolites Parkinson 1811LT64 [incl. Checchiaites Sorrentino 1935LT64, Eoalveolinella Silvestri 1928LT64, Flosculina Stache in Schwager 1883LT64Miliolites de Montfort 1808 non Lamarck 1804LT64, Oryzaria Defrance in Bronn 1825LT64, Semiflosculina Doncieux 1905LT64]

*Type species of generic name indicated

References

[B-F08] BouDagher-Fadel, M. K. 2008. The Cenozoic larger benthic foraminifera: the Palaeogene. Developments in Palaeontology and Stratigraphy 21: 297–418.

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3rd ed. Harvard University Press: Cambridge (Massachusetts).

[HW93] Hart, M. B., & C. L. Williams. 1993. Protozoa. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 43–70. Chapman & Hall: London.

[HS02] Höfling, R., & R. W. Scott. 2002. Early and Mid-Cretaceous buildups. SEPM Special Publication 72: 521–548.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[M12] McGowran, B. 2012. Cenozoic environmental shifts and foraminiferal evolution. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 937–965. Springer.

[MP14] Merle, D., J.-M. Pacaud, G. Métais, A. Bartolini, R. A. Lashari, I. A. Brohi, S. H. Solangi, L. Marivaux & J.-L. Welcomme. 2014. Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography. Zootaxa 3826 (1): 101–138.

[PHT13] Pawlowski, J., M. Holzmann & J. Tyszka. 2013. New supraordinal classification of Foraminifera: molecules meet morphology. Marine Micropalaeontology 100: 1–10.

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