Belongs within: Panheteroptera.
Contains: Mezirinae.
The Aradoidea are a group of flattened bugs, many species of which are found on or under bark of dead or dying trees (Grimaldi & Engel 2005). When wings are present, the sides of the abdomen extend outside the margins of the wings at rest. Most living species are included in the family Aradidae with the exception of the Termitaphididae, a circumtropical group of blind, wingless species that live in termite nests (Carver et al. 1991). A separate family has also been accorded Kobdocoris aradinus of the earliest Cretaceous of Mongolia (Ross & Jarzembowski 1993).
Synapomorphies (from Shcherbakov & Popov 2002): Body strongly flattened dorsoventrally; extremely long stylets coiled within head; abdominal trichobothria absent.
<==Aradoidea [Aradomorpha]
|--KobdocorisRJ93 [KobdocoridaeSP02]
| `--K. aradinus Popov 1986RJ93
|--Termitaradus [Termitaphididae]GE05
| |--T. australiensisCGW91
| `--T. guianaeYR12
`--AradidaeGE05
| i. s.: Clavaptera ornata Kirman 1985Lar02
| Leuraptera yakasi Heiss 1990Lar02
| Neocarventus uncus Kirman 1989Lar02
| Archearadus burmensisGE05
| Odonia Bergroth 1889B89
| `--*O. truncata [=Aradus truncatus]B89
| ArtabanusB89
| |--A. atkinsoni Bergroth 1889B89
| |--A. breviscutum Bergroth 1889B89
| `--A. geniculatusB89
| BrachyrrhynchusB89
| |--B. feanus Bergroth 1889B89
| |--B. membranaceus [incl. B. albipennis, B. orientalis]B89
| |--B. montanus Bergroth 1889B89
| |--B. scabrosusB89
| |--B. setosusB89
| |--B. tagalicus Stål 1870B89
| `--B. triangulus Bergroth 1889B89
| LlaimacorisWEE74
| Emydocoris Usinger 1944D56
| |--E. montanus Drake 1956D56
| `--E. testidinatusD56
| Reeceicus Drake 1956D56
| `--*R. saileri Drake 1956D56
| Glyptocoris verus Drake 1956D56
| AglaocorisD56
| |--A. cubanus Drake 1956D56
| |--A. comes Drake 1956D56
| `--A. nataliiD56
| Asterocoris (Peggicoris Drake 1956)D56
| `--A. (*P.) zeteki Drake 1956D56
| Mastigocoris insularis Kormilev 1983JG19
| AcanthiaL49
| |--A. lectulariaL49
| |--A. littoralisLat02
| |--A. saltatoriaG20
| `--A. zosteraeLat02
| Piestosoma flavomaculata [=P. flavomaculatum]L49
|--Prosympiestus [Prosympiestinae]CGW91
|--MezirinaeCGW91
|--Isodermus [Isoderminae]CGW91
| `--I. planusCGW91
|--ChinamyersiinaeCGW91
| |--Kumaressa scutellataCGW91
| `--TretocorisCGW91
|--CalisiusLar02 [CalisiinaeCGW91]
| |--C. interveniensK08
| |--C. pacificus Kirkaldy 1908K08
| `--C. zealandicus Pendergrast 1968Lar02
|--CarventinaeCGW91
| |--CarventusCGW91
| |--GlyptoapteraCGW91
| |--ProbatocepsCGW91
| `--Acaraptera waipouensis Heiss 1990CGW91, Lar02
|--AneurusLar02 [AneurinaeCGW91]
| | i. s.: A. prominens Pendergrast 1965Lar02
| `--A. (Aneurodellus)Lar02
| |--A. (A.) brevipennis Heiss 1998Lar02
| |--A. (A.) maoricus Heiss 1998Lar02
| `--A. (A.) zealandensis Heiss 1998Lar02
`--AradusTKM93 [AradinaeCGW91]
| i. s.: A. abnormis Bergroth 1889B89
| A. betulaeG20
| A. candidatus Bergroth 1889B89
| A. creticusYR12
| A. flavicornisB89
| A. frigidus Kiritshenko 1913TKM93
| A. lugubris Boisduval 1835B35
| A. nicholasi Popov 1989RJ93
| A. variusB49
`--A. (Quilnus)H01
|--A. (Q.) brevirostris Horváth 1901H01
`--A. (Q.) nigerH01
*Type species of generic name indicated
References
[B49] Baker, E. W. 1949. A review of the mites of the family Cheyletidae in the United States National Museum. Proceedings of the United States National Museum 99 (3238): 267–320.
[B89] Bergroth, E. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XXII.—E. Bergrothii Commentarius de Aradidis in Burma et Tenasserim a L. Fea collectis. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 730–739.
[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d’Urville. Faune entomologique de l’océan Pacifique, avec l’illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.
[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).
[D56] Drake, C. J. 1956. New Neotropical genera and species of apterous aradids (Hemiptera). Journal of the Washington Academy of Sciences 46 (10): 322–327.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.
[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.
[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.
[Lar02] Larivière, M.-C. 2002. Primary types of Heteroptera in the New Zealand Arthropod Collection: Pentatomomorpha described from New Zealand before the year 2000. New Zealand Entomologist 25: 79–85.
[Lat02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.
[L49] Lucas, H. 1849. Exploration Scientifique de l’Algérie pendant les années 1840, 1841, 1842 publiée par ordre du gouvernement et avec le concours d’une commission académique. Sciences physiques. Zoologie. II. Histoire naturelle des animaux articulés. Troisième partie. Insectes (suite). Imprimerie Nationale: Paris.
[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.
[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815+Heteroptera Latreille, 1810). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 143–157. Kluwer Academic Publishers: Dordrecht.
[TKM93] Thaler, K., B. Knoflach & E. Meyer. 1993. Fragmenta faunistica Tirolensia—X. Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 80: 311–325.
[WEE74] Woodward, T. E., V. F. Eastop & J. W. Evans. 1974. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 52–57. Melbourne University Press.
[YR12] Yao, Y., D. Ren, D. A. Rider & W. Cai. 2012. Phylogeny of the infraorder Pentatomomorpha based on fossil and extant morphology, with description of a new fossil family from China. PLoS One 7 (5): e37289.