Archidermaptera

 The Late Jurassic Microdiplatys campodeiformis, from Grimaldi & Engel (2005).

Belongs within: Forficulida.

The Archidermaptera are a basal, fossil group within the Dermaptera (earwigs) known from the late Triassic to the early Cretaceous. The group is defined solely by plesiomorphic characters and may be paraphyletic. Archidermapterans had the shortened, hardened forewings (tegmina) of modern earwigs, but the tegmina still retained some veins and the cerci had not yet been modified into the forceps characteristic of modern earwigs.

Characters (from Grimaldi & Engel 2005): Tegmina veined; ovipositor well-developed; cerci long and multi-segmented; tarsi five-segmented.

<==Archidermaptera
    |--TuranoviidaeGE05
    |--Dermapteridae [Dermapterinae]GE05
    |--SinopalaeodermatidaeZ02
    |    |--Sinopalaeodermata Zhang 2002Z02
    |    |    `--*S. neimonggolensis Zhang 2002Z02
    |    `--Jurassimedeola Zhang 2002Z02
    |         `--*J. orientalis Zhang 2002Z02
    |--LongicerciatidaeZ02
    |    |--Sinostaphylina nanligezhuangensis Hong & Wang 1990Z02
    |    `--LongicerciataZ02
    |         |--L. mesozoica Zhang 1994NW03
    |         `--L. rumpens Zhang 1994Z02
    `--Protodiplatyidae [Protodiplateinae, Protodiplateoidea, Protodiplateomorpha, Protodiplatidae]Z02
         |--Baseopsis forficulinaS02
         |--Asiodiplatys speciosusS02
         |--MicrodiplatysGE05
         |    |--M. campodeiformisGE05
         |    `--M. perfectus Vishniakova 1985Z02
         `--ProtodiplatysZ02
              |--*P. fortis Martynov 1925Z02
              |--P. gracilis Vishniakova 1980K03
              `--P. mongoliensis Vishniakova 1986RJ93

*Type species of generic name indicated

References

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[K03] Kluge, N. J. 2003. About evolution and homology of genital appendages of insects. Trudy Russkogo Entomologicheskogo Obshestva [Proceedings of the Russian Entomological Society] 74: 3–16.

[NW03] Nel, A., A. Waller, V. Albouy, J.-J. Menier & G. de Ploëg. 2003. New fossil earwigs from the lowermost Eocene amber of Paris Basin (France) (Insecta, Dermaptera, family incertae sedis). Geodiversitas 25 (1): 119–129.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[S02] Shcherbakov, D. E. 2002. Order Forficulida Latreille, 1810. The earwigs and protelytropterans (=Dermaptera DeGeer, 1773 +Protelytroptera Tillyard, 1931). In: Rasnitsyn, A. P.,  & D. L. J. Quicke (eds) History of Insects pp. 288–291. Kluwer Academic Publishers: Dordrecht.

[Z02] Zhang, J.-F. 2002. The most primitive earwigs (Archidermaptera, Dermaptera, Insecta) from the Upper Jurassic of Nei Monggol Autonomous Region, northeastern China. Acta Micropalaeontologica Sinica 19 (4): 348–362.

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