Belongs within: Boletales.
Contains: Leccinum, Boletus.
The Boletaceae are a clade of fungi commonly producing fleshy fruiting bodies with poroid hymenophores.
<==Boletaceae [Boletineae, Chamonixiaceae, Strobilomycetaceae, Xerocomaceae]
|--Chalciporus Bataille 1908BB02b, KC01
| |--C. amarellusBB02b
| `--C. piperatusBB02b
`--+--BoletusBB02b
`--+--Retiboletus Binder & Bresinsky 2002BB02b
| |--*R. ornatipes (Peck) Binder & Bresinsky 2002 [=Boletus ornatipes Peck 1878]BB02b
| |--R. flavoniger (Halling, von Mueller & Gómez) Binder & Bresinsky 2002 (see below for synonymy)BB02b
| |--R. griseus (Frost) Binder & Bresinsky 2002 [=Boletus griseus Frost in Peck 1878]BB02b
| |--R. nigerrimus (Heim) Binder & Bresinsky 2002 [=Boletus nigerrimus Heiim 1963, Tylopilus nigerrimus]BB02b
| `--R. retipes (Berkeley & Curtis) Binder & Bresinsky 2002 [=Boletus retipes Berkeley & Curtis 1872]BB02b
`--+--Fistulinella Henn. 1901BB02b, KC01 [incl. Ixechinus Heim 1968KC01, Mucilopilus Wolfe 1979KC01]
| `--F. viscidaBB02b
|--Tylopilus Karst. 1881BB02b, KC01 (see below for synonymy)
| | i. s.: T. chromapes [=Leccinum chromapes]BB00
| | ‘Boletus’ cyanopusW01
| | T. indecisusBB02b
| | T. maculatusW01
| | T. rubrobrunneusKC01
| | T. vinaceipallidusW01
| |--T. alboaterBB00
| `--+--T. felleusBB02b
| `--T. ferrugineusBB00
`--+--Xerocomus Quél. 1887 (nom. cons.)BB02b, KC01
| | i. s.: X. astraeicolaW01
| | X. (sect. *Pseudoboleti) badius [=Boletus badius]W01
| | X. chrysenteronHB02
| |--+--X. depilatus (Redeuilh) Binder & Besl 2000 (see below for synonymy)BB00
| | `--X. impolitus [=Boletus impolitus]BB00
| `--+--X. hortonii (Smith & Thiers) Binder & Besl 2000 (see below for synonymy)BB00
| `--+--*X. subtomentosusBB00
| `--X. illudensBB00
`--Phylloporus Quél. 1888BB02b, KC01
|--P. bellusW01
|--P. borgorienseW01
|--P. foliiporusW01
|--P. leucomycelinusW01
|--P. pelletieriBB02b
`--P. rhodoxanthusBB02b
Boletaceae incertae sedis:
Gastroboletus Lohwag 1926KC01
|--*G. boedijniiW01
`--G. turbinatusBB02a
LeccinumKC01
Chamonixia Rolland 1899KC01
|--C. caespitosaBB02a
`--C. mucosaW01
Boletellus Murrill 1909 (see below for synonymy)KC01
|--B. ananasW01
|--B. chrysenteroidesW01
|--B. emodensisW01
|--B. longicollisW01
|--B. obscureococcineusW01
`--B. projectellusJK06
Pulveroboletus Murrill 1909KC01
|--*P. ravenelliiW01
|--P. friansW01
|--P. icterinusW01
`--P. retipesW01
Austroboletus (Corner) Wolfe 1980KC01
|--A. dictyopusW01
|--A. malaccensis [=Phylloporus malaccensis, Porphyrellus malaccensis]W01
|--A. mucosus [=Boletus mucosus]W01
|--A. rarusW01
|--A. rostrupiiW01
`--A. rubricolorW01
Xanthoconium Singer 1944KC01
Rubinoboletus Pilát & Dermek 1969W01, KC01
`--*R. rubinus [=Boletus rubinus]W01
Aureoboletus Pouzar 1957KC01
|--‘Boletus’ thibetanusW01
`--‘Boletus’ umbilicatus [incl. B. corrugatus]W01
Pseudoboletus Sutara 1991 [=Xerocomus sect. Parasitici]W01
|--‘Boletus’ astraeicolopsisW01
`--‘Xerocomus’ parasiticusW01
Afroboletus Pegler & Young 1981KC01
Strobilomyces Berk. 1851 [=Eriocorys Quél. 1886]KC01
|--S. annulatusW01
|--S. floccopusSS09
|--S. foveatusW01
|--S. mirandusW01
|--S. nigricansW01
|--S. polypyramisW01
`--S. velutipesW01
Boletochaete Singer 1944KC01
Buchwaldoboletus Pilát 1969KC01
|--B. spectabilisW01
`--B. sulphureusW01
Gastroleccinum Thiers 1989KC01
Gastrotylopilus Li & Watling 1999KC01
Heimiella Boedijn 1951KC01 [incl. Boletus subg. Punctispora Corner 1974W01]
|--H. anguiformisW01
|--H. kinabulensisW01
|--H. mandarinaW01
|--‘Boletus (subg. *Punctispora)’ punctisporus Corner 1974W01
|--H. retisporaW01
`--H. subretisporaW01
Paxillogaster Horak 1966KC01
Phylloboletellus Singer 1952KC01
Royoungia Castellano, Trappe & Malajczuk 1992KC01
Setogyroporus Heinem. & Rammeloo 1982KC01
Singeromyces Moser 1966KC01
Sinoboletus Zang 1992KC01
Tubosaeta Horak 1967KC01
Veloporphyrellus Gómez & Singer 1984KC01
Boletellus Murrill 1909 [incl. Boletogaster Lohwag 1926, Frostiella Murrill 1942 (n. n.), Strobilofungus Lloyd 1915]KC01
Retiboletus flavoniger (Halling, von Mueller & Gómez) Binder & Bresinsky 2002 [=Boletus flavoniger Halling, von Mueller & Gómez 1999]BB02b
Tylopilus Karst. 1881BB02b, KC01 [=Rhodoporus Quél. ex Bataille 1908KC01; incl. Leucogyroporus Snell 1942KC01, Phaeoporus Bataille 1908 non Schröt. 1888KC01, Porphyrellus Gilbert 1931KC01, Rhodobolites Beck 1923KC01]
Xerocomus depilatus (Redeuilh) Binder & Besl 2000 [=Boletus depilatus Redeuilh 1985, Leccinum depilatum (Redeuilh) Šutara 1989]BB00
Xerocomus hortonii (Smith & Thiers) Binder & Besl 2000 [=Boletus hortonii Smith & Thiers 1971, Leccinum hortonii (Smith & Thiers) Hongo & Nagasawa 1978; incl. B. subglabripes ssp. corrugis Peck 1889]BB00
*Type species of generic name indicated
References
[BB00] Binder, M., & H. Besl. 2000. 28S rDNA sequence data and chemotaxonomical analyses on the generic concept of Leccinum (Boletales). In: A. M. Bresadola (ed.) Micologia 2000 pp. 75–86. AMB, Fundacione Centro Studi Micologici: Trento (Italy).
[BB02a] Binder, M., & A. Bresinsky. 2002a. Derivation of a polymorphic lineage of Gasteromycetes from boletoid ancestors. Mycologia 94 (1): 85–98.
[BB02b] Binder, M., & A. Bresinsky. 2002b. Retiboletus, a new genus for a species-complex in the Boletaceae producing retipolides. Feddes Repertorium 113 (1–2): 30–40.
[HB02] Hibbett, D. S., & M. Binder. 2002. Evolution of complex fruiting-body morphologies in homobasidiomycetes. Proceedings of the Royal Society of London Series B—Biological Sciences 269: 1963–1969.
[JK06] James, T. Y., F. Kauff, C. L. Schoch, P. B. Matheny, V. Hofstetter, C. J. Cox, G. Celio, C. Gueidan, E. Fraker, J. Miadlikowska, H. T. Lumbsch, A. Rauhut, V. Reeb, A. E. Arnold, A. Amtoft, J. E. Stajich, K. Hosaka, G.-H. Sung, D. Johnson, B. O’Rourke, M. Crockett, M. Binder, J. M. Curtis, J. C. Slot, Z. Wang, A. W. Wilson, A. Schüßler, J. E. Longcore, K. O’Donnell, S. Mozley-Standridge, D. Porter, P. M. Letcher, M. J. Powell, J. W. Taylor, M. M. White, G. W. Griffith, D. R. Davies, R. A. Humber, J. B. Morton, J. Sugiyama, A. Y. Rossman, J. D. Rogers, D. H. Pfister, D. Hewitt, K. Hansen, S. Hambleton, R. A. Shoemaker, J. Kohlmeyer, B. Volkmann-Kohlmeyer, R. A. Spotts, M. Serdani, P. W. Crous, K. W. Hughes, K. Matsuura, E. Langer, G. Langer, W. A. Untereiner, R. Lücking, B. Büdel, D. M. Geiser, A. Aptroot, P. Diederich, I. Schmitt, M. Schultz, R. Yahr, D. S. Hibbett, F. Lutzoni, D. J. McLaughlin, J. W. Spatafora & R. Vilgalys. 2006. Reconstructing the early evolution of Fungi using a six-gene phylogeny. Nature 443: 818–822.
[KC01] Kirk, P. M., P. F. Cannon, J. C. David & J. A. Stalpers. 2001. Ainsworth & Bisby’s Dictionary of the Fungi 9th ed. CAB International: Wallingford (UK).
[SS09] Schoch, C. L., G.-H. Sung, F. López-Giráldez, J. P. Townsend, J. Miadlikowska, V. Hofstetter, B. Robbertse, P. B. Matheny, F. Kauff, Z. Wang, C. Gueidan, R. M. Andrie, K. Trippe, L. M. Ciufetti, A. Wynns, E. Fraker, B. P. Hodkinson, G. Bonito, J. Z. Groenewald, M. Arzanlou, G. S. de Hoog, P. W. Crous, D. Hewitt, D. H. Pfister, K. Peterson, M. Gryzenhout, M. J. Wingfield, A. Aptroot, S.-O. Suh, M. Blackwell, D. M. Hillis, G. W. Griffith, L. A. Castlebury, A. Y. Rossman, H. T. Lumbsch, R. Lücking, B. Büdel, A. Rauhut, P. Diederich, D. Ertz, D. M. Geiser, K. Hosaka, P. Inderbitzin, J. Kohlmeyer, B. Volkmann-Kohlmeyer, L. Mostert, K. O’Donnell, H. Sipman, J. D. Rogers, R. A. Shoemaker, J. Sugiyama, R. C. Summerbell, W. Untereiner, P. R. Johnston, S. Stenroos, A. Zuccaro, P. S. Dyer, P. D. Crittenden, M. S. Cole, K. Hansen, J. M. Trappe, R. Yahr, F. Lutzoni & J. W. Spatafora. 2009. The Ascomycota tree of life: a phylum-wide phylogeny clarifies the origin and evolution of fundamental reproductive and ecological traits. Systematic Biology 58 (2): 224–239.
[W01] Watling, R. 2001. The relationships and possible distributional patterns of boletes in south-east Asia. Mycological Research 105 (12): 1440–1448.