Belongs within: Curculioninae.
Contains: Acamptini, Pentarthrini, Dryotribini, Cossonini, Stenoscelidina, Rhyncolina.
The Cossoninae are a group of elongate-bodied weevils that are borers in dead or dying wood (Marvaldi & Lanteri 2005).
Characters (from Marvaldi & Lanteri 2005): Body elongate, parallel-sided, depressed. Rostrum often shorter than prothorax. All legs usually with uncinate tibiae; front tibiae with grooming device consisting of long comb of setae on apical third to half; hind tibiae lacking apical comb of setae. Tarsite 3 as wide as tarsite 2.
<==Cossoninae [Cossonides] |--AraucariiniM94 | |--AraucariusMPJ18 | | |--A. majorMPJ18 | | `--A. minorMPJ18 | `--+--CoptocorynusMPJ18 | `--Xenocnema Wollaston 1873MPJ18, M94 | `--*X. spinipes Wollaston 1873M94 |--+--CossoniniMPJ18 | `--OnycholipiniL20 | |--Onycholips bifurcatus Wollaston 1861L20 | |--PseudostenocelisMPJ18 | |--Trichacorynus Blatchley 1916A02 | |--Pselactus Broun 1886MPJ18, A02 [incl. Codiosoma Bedel 1885 non Kirby 1874A02, Phloeophagia Aurivillius 1924A02] | | `--P. spadix (Herbst 1795)HF03 | `--Pseudopentarthrum Wollaston 1873 (see below for synonymy)A02 | `--P. robustumA02 `--RhyncoliniM94 | i. s.: Phloeophagosoma Wollaston 1873M94 | |--*P. minutum Wollaston 1873M94 | |--P. aesculiM94 | |--P. tenuisM94 | `--P. thoracicum Wollaston 1874M94 |--StenoscelidinaL20 |--Phloeophagus Schoenherr 1838A02 [PhloeophaginaL20] |--RhyncolinaMPJ18 |--ChoerorhinusHF03 [ChoerorhininaL20] | `--C. squalidus Fairmaire 1858L20 `--Pseudomimina [Aphyllurini]L20 |--Aphyllura brenskei Reitter 1884L20 `--PseudomimusHF03 |--P. avocadi Folwaczny 1973HF03 `--P. burgeoni Hustache 1934L20 Cossoninae incertae sedis: Phoenomerus exilis Pascoe 1872M86 Notiomimetes pascoei Woll. 1873M86 CossonideusM86 |--C. lineola (Fabricius 1775) [=Curculio lineola]Z93 `--C. pascoei Woll. 1873M86 Halorhynchus caecus Woll. 1873M86 Thaumastophasis oculatus Woll. 1873M86 Stereomimetes crassicornis Woll. 1873M86 NotiosomusM86 |--N. australis Woll. 1873M86 |--N. congener Woll. 1873M86 `--N. major Woll. 1873M86 Homalotrogus arctatus Pascoe 1885M86 Aphanocorynes depressus Woll. 1873M86 PentamimusM86 |--P. canaliculatus Woll. 1873M86 |--P. rhyncholiformis Woll. 1873M86 `--P. suffusus Woll. 1873M86 OrthotemnusL20 |--O. disparilis Pascoe 1885M86 `--O. filiformis Champion 1914L20 StereoborusM86 |--S. inductus Pascoe 1885M86 |--S. interstitialis Lea 1896L96 |--S. laporteae Lea 1896L96 `--S. macleayi Lea 1896L96 AphyodaZ93 EiratusM94 ExeiratusM94 Euophryum Broun 1909L20, A02 `--E. confine (Broun 1881)L20 Caulotrupis lucifugus Wollaston 1854L20 AcamptiniZ93 PentarthriniM94 Pseudapotrepus [Pseudapotrepini]L20 `--P. insularis Hustache 1932L20 CryptommatiniL20 TapiromiminiL20 BrachytemnusGC-P14 [NeumatoriniL20] `--B. porcatus Germar 1824HF03 DryotribiniL20 Nesiobius [Nesiobiini]L20 `--N. horridus (Wollaston 1871)L20 Acanthinomerus [Acanthinomerini]L20 `--A. conicollis (Wollaston 1861)L20 Microxylobius [Microxylobiini]L20 `--M. lucifugus Wollaston 1861L20 HimatininiL20 AllomorphiniL20 Hexarthroides Lea 1896L96 `--*H. punctulatum Lea 1896L96 Proeces Schoenherr 1838 (see below for synonymy)A02 `--P. depressus (Boheman 1838)A02
Proeces Schoenherr 1838 [incl. Borborhynchus Richard 1957, Eucoptus Wollaston 1873, Stenotis Wollaston 1854; Proecini]A02
Pseudopentarthrum Wollaston 1873 [incl. Micropentarthrum Champion 1909, Neopentarthrum Mutchler 1925, Pentarthrinus Casey 1892, Phloeophagomorphus Wollaston 1873, Stenotylus Marshall 1933]A02
*Type species of generic name indicated
References
[A02] Anderson, R. S. 2002. Curculionidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 722–815. CRC Press.
[GC-P14] Gillett, C. P. D. T., A. Crampton-Platt, M. J. T. N. Timmermans, B. H. Jordal, B. C. Emerson & A. P. Vogler. 2014. Bulk de novo mitogenome assembly from pooled total DNA elucidates the phylogeny of weevils (Coleoptera: Curculionoidea). Molecular Biology and Evolution 31 (8): 2223–2237.
[HF03] Halperin, J., & J. Fremuth. 2003. Contribution to the knowledge of Curculionoidea (Coleoptera) and their host plants in Israel. Zoology in the Middle East 29: 93–100.
[L96] Lea, A. M. 1896. Descriptions of new species of Australian Coleoptera. Part III. Proceedings of the Linnean Society of New South Wales 21 (3): 284–319.
[L20] Legalov, A. A. 2020. Annotated key to weevils of the world. Part 4. Subfamilies Erirhininae, Dryophthorinae and Cossoninae (Curculionidae). Ukrainian Journal of Ecology 10 (2): 319–331.
[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part V. Proceedings of the Linnean Society of New South Wales, series 2, 1 (3): 585–686.
[M94] May, B. M. 1994. An introduction to the immature stages of Australian Curculionoidea. In: Zimmerman, E. C. Australian Weevils (Coleoptera: Curculionoidea) vol. 2. Brentidae, Eurhynchidae, Apionidae and a chapter on immature stages by Brenda May pp. 365–728. CSIRO Australia.
[MPJ18] Mugu, S., D. Pistone & B. H. Jordal. 2018. New molecular markers resolve the phylogenetic position of the enigmatic wood-boring weevils Platypodinae (Coleoptera: Curculionidae). Arthropod Systematics and Phylogeny 76 (1): 45–58.
[Z93] Zimmerman, E. C. 1993. Australian Weevils (Coleoptera: Curculionoidea) vol. 3. Nanophyidae, Rhynchophoridae, Erirhinidae, Curculionidae: Amycterinae, literature consulted. CSIRO Australia.