Belongs within: Coccinelloidea.
Contains: Endomychinae.
The Endomychidae are a group of often brightly coloured fungus-feeding beetles. Larvae often bear lateral tergal processes on most segments; in the species Encymon immaculatus, these processes are spinose and may break off to deter predators (Lawrence & Britton 1991). Members of the genus Trochoideus have remarkable antennae with just five segments, the last two of which are greatly enlarged and appear fused (Skelley & Leschen 2002).
Characters (from Lawrence & Britton 1991): Broadly ovate to elongate and globose to moderately flattened, glabrous or clothed with decumbent and erect hairs, often brightly patterned with red or yellow and black. Frontoclypeal suture present; antennae moderately long, usually with relatively loose club; maxillary palps fusiform, labial palps may be securiform; fore coxae globular, trochantins concealed, fore coxal cavities externally open or closed, internally closed; mid coxal cavities moderately to widely separated, laterally open or closed; tarsi trimerous or tetramerous, second tarsal segment commonly lobed, extending beneath reduced third segment; tarsal claws simple; abdomen with five or six ventrites, none connate, femoral lines on first ventrite usually absent. Larva usually oblong to ovate (rarely elongate), slightly to strongly flattened, usually with lateral tergal processes on all thoracic and most abdominal segments and only rarely with short urogomphi; antennae often long (with segment 2 elongate and 1 and 3 reduced), their sockets well separated from mandibular articulations; head often broad at base, strongly narrowed towards mouth-parts; mala obtuse.
<==Endomychidae [Endomychides]
|--+--Trochoideus Westwood 1833SL02 [PleganophorinaeRS15]
| | |--T. boliviensisRS15
| | |--T. cruciatus (Dalm. 1835) [=Pausus cruciatus]LC40
| | |--T. desjardinsiLB91
| | `--T. goudotiRS15
| `-+--LeiestinaeRS15
| | |--Stethorhanis Blaisdell 1931SL02
| | |--Rhanidea Strohecker 1953 [incl. Rhanis LeConte 1854 non Heyden 1837]SL02
| | | `--R. unicolorSL02
| | `--Phymaphora Newman 1838SL02
| | |--P. californicaSL02
| | `--P. pulchellaSL02
| `--Merophysiinae [Holoparamecinae, Merophysiidae]RS15
| |--MerophysiaRD77
| |--Holoparamecus Curtis 1833 [incl. Calyptobium Aubé 1843]SL02
| | |--H. depressusLB91
| | |--H. dispar Belon 1885B89
| | `--H. kunzeiMW15
| `--Lycoperdinella subcaecaRS15
`--+--Cyclotominae [Cyclotomidae]RS15
| |--Cyclotoma Mulsant 1851RS15
| | `--C. cingalensisRS15
| |--MeilichiusRS15
| |--BolbomorphusRS15
| `--EucteanusRS15
`--+--EndomychinaeRS15
`--+--EpipocinaeRS15
| | i. s.: Hadromychus Bousquet & Leschen 2002SL02
| | `--H. chandleriSL02
| |--Epipocus Germar 1943RS15, SL02
| `--+--AnidrytusRS15
| `--Epopterus Chevrolat 1844RS15, SL02
| `--E. testudinariusRS15
`--LycoperdininaeRS15
|--Mycetina Mulsant 1846RS15, SL02
| |--M. amabilisRS15
| `--M. horniRS15
`--+--Lycoperdina Latreille 1907RS15, SL02
| |--L. bovistaeG20
| |--L. ferrugineaSL02
| `--L. immaculataR26
`--+--+--Acinaces laceratusRS15
| `--+--BeccariolaRS15
| `--Corynomalus [incl. Amphix]RS15
| |--C. vestitusRS15
| `--+--C. laevigatusRS15
| `--C. tarsatusRS15
`--+--Aphorista Gorham 1873RS15, SL02
| |--A. morosaV99
| `--A. vittataRS15
`--+--+--TrycherusRS15
| `--AmphisternusRS15
`--+--EncymonRS15
| |--E. clavicornisC70
| |--E. gorhamiRS15
| `--E. immaculatusLB91
`--+--EumorphusRS15
| |--E. immarginatusG20
| `--E. quadriguttatus (Illiger 1800)B14
`--+--Indalmus lineelusRS15
`--AncylopusRS15
|--A. bisignatusG01b
|--A. fuscipennisG00
|--A. melanocephalusG01b
|--A. nigrofuscus Gorham 1900G01a
`--A. unicolorG01b
Endomychidae incertae sedis:
DisploteraLB91
DapsaC01
|--D. barbara Lucas 1846E12
|--D. (Phylira) horvathi Csiki 1901C01
|--D. limbataC01
|--D. nigripennisC01
`--D. trimaculataC01
Phymaphoroides antennatusP92
HylaiaP92
LiesthesP92
CholovoceraRS15
Xenomycetes Horn 1880 [Xenomycetinae]SL02
|--X. laversiSL02
`--X. morrisoniSL02
DanascelinaeRS15
CholovoceridaRS15
*Type species of generic name indicated
References
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[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.
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[SL02] Skelley, P. E., & R. A. B. Leschen. 2002. Endomychidae Leach 1815. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 366–370. CRC Press.
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