Belongs within: Polypodiaceae.
Contains: Grammitis.
The Grammitidaceae are a mostly epiphytic family of ferns. Many species are found in montane cloud forests. Members typically bear scaled rhizomes and rounded, abaxial sori.
Characters (A. R. Smith): Plants perennial, mostly small, on rock or commonly epiphytic [rarely terrestrial]. Stems long- to short-creeping or suberect, usually unbranched, bearing scales [rarely scales absent], solenostelic (having phloem on both sides of xylem) to dictyostelic (having complex nets of xylem). Leaves erect, arching, or pendent, monomorphic [rarely with specialized fertile areas], less than 50 cm [rarely longer], usually scaleless throughout. Petioles often dark-colored and wiry, commonly terete, usually less than 2 mm diam., articulate or not articulate, with 1 or 2 vascular strands. Blades simple and entire to commonly pinnatifid or 1-pinnate, rarely 2-pinnate or more divided, glabrous or commonly bearing hairs, especially on petioles and rachises; hairs tan to dark reddish brown [or transparent], unicellular to multicellular; rachises often dark-colored, not grooved adaxially. Veins free [to anastomosing in simple patterns]; hydathodes present or absent, sometimes obscured by lime dots adaxially. Sori abaxial on veins, round to oblong [occasionally elongate]; paraphyses present or absent, these glandular or hairlike; sporangia with stalk of 1 row of cells; indusia absent. Spores greenish, tetrahedral-globose, trilete, surface commonly papillate. Gametophytes greenish, borne aboveground, ribbon-shaped, sometimes bearing multicellular gemmae.
Grammitidaceae
| i. s.: LomaphlebiaP03
| LuismaP03
| AcrosorusP03
| ThemeliumP03
|--‘Terpsichore’ eggersiiSP09
`--+--+--+--Adenophorus pinnatifidusSP09
| | `--‘Grammitis’ tenellaSP09
| `--+--CochlidiumSP09 [incl. XiphopterisP03]
| | |--‘Xiphopteris’ hieronymusiiP03
| | |--C. seminudumSP09
| | |--C. serrulatum [=*Xiphopteris serrulata]P03
| | `--‘Xiphopteris’ sikkimensis (Hieronymus) Copel. 1947 [=Polypodium sikkimense Hieronymus 1905]I88
| `--Enterosora parietinaSP09
`--+--+--‘Lellingeria’ schenckiiSP09
| `--‘Terpsichore’ senilisSP09
`--+--CeradeniaSP09
| |--C. kalbreyeriSP09
| `--C. pilipesSP09
`--+--TerpsichoreSP09
| | i. s.: T. elasticaP03
| | T. kirkiiP03
| |--T. anfractuosaSP09
| `--+--T. semihirsutaSP09
| `--+--MelpomeneSP09
| | |--M. flabelliformisP03
| | `--M. moniliformisSP09
| `--LellingeriaSP09
| |--L. apiculataSP09
| |--L. hirsutaSP09
| |--L. oosoraP03
| |--L. saffordiiP03
| `--L. subcoriaceaP03
`--+--+--Chrysogrammitis musgravianaSP09
| `--MicropolypodiumSP09
| |--M. hyalinumSP09
| `--M. taenifoliumSP09
`--+--+--Calymmodon gracilisSP09
| `--+--‘Ctenopteris’ repandulaSP09
| `--Scleroglossum sulcatumSP09
`--+--+--‘Ctenopteris’ lasiostipesSP09
| `--GrammitisSP09
`--+--+--CtenopterisSP09
| | |--*C. venulosa [=Prosaptia venulosa]P03
| | |--C. blechnoidesP03
| | |--C. curtisiiP03
| | |--C. mollicomaP03
| | |--C. nutansP03
| | |--C. subfalcata (Blume) Kunze 1848 [=Polypodium subfalcatum Blume 1828]I88
| | `--C. whartonianaP03
| `--ProsaptiaSP09
| |--P. contiguaSP09
| `--P. obliquataSP09
`--+--‘Grammitis’ conjunctisoraSP09
`--+--‘Grammitis’ hookeriSP09 (see below for synonymy)
`--‘Grammitis’ parvaSP09
‘Grammitis’ hookeriSP09 [=Polypodium hookeri Brackenridge 1854W16, P. setigerum Hook. & Arn. 1832 non Blume 1828W16]
*Type species of generic name indicated
References
[I88] Iwatsuki, K. 1988. An enumeration of the pteridophytes of Nepal. In: Ohba, H., & S. B. Malla (eds) The Himalayan Plants vol. 1. The University Museum, University of Tokyo, Bulletin 31: 231–339.
[P03] Parris, B. S. 2003. The distribution of Grammitidaceae (Filicales) inside and outside Malesia. Telopea 10 (1): 451–466.
[SP09] Schuettpelz, E., & K. M. Pryer. 2009. Evidence for a Cenozoic radiation of ferns in an angiosperm-dominated canopy. Proceedings of the National Academy of Sciences of the USA 106 (27): 11200–11205.
[W16] Watts, W. W. 1916. Two Lord Howe Island Polypodia. Journal and Proceedings of the Royal Society of New South Wales 49 (3): 385–388.