Belongs within: Panheteroptera.
The Leptopodomorpha are a group of large-eyed, predatory bugs most commonly found on the margins of ponds and streams. They are united by adaptations to a side-by-side copulatory position, with the male abdomen possessing an apparatus for grasping the female (Schuh & Polhemus 1980). The fore wing is elytriform or bears open cells on the apical area in Leptopodoidea whereas members of the Saldoidea have four or five closed cells on the membrane of the fore wing (Carver et al. 1991). Within the Leptopodoidea, the Indo-Pacific Omaniidae are beetle-like and flightless, with hemelytra lacking a membrane and meeting in a straight line (Carver et al. 1991).
<==Leptopodomorpha [Acanthioideae, Leptopinae]
| i. s.: BritannicolaGE05
| Helenosaldula aberransSP80
| Archegocimicidae [Archaegocimicidae, Diatilidae, Eonabidae]GE05
| |--Saldonia sibiricaSP02
| `--Sondalia kovalevi Popov 1988RJ93
|--LeptopodoideaSP80
| |--OmaniidaeSP80
| | |--OmaniaSP80
| | | |--O. coleoptrataSP80
| | | `--O. marksaeWEE70
| | `--CorallocorisSP80
| | |--C. marksaeCGW91
| | `--C. nauruensisSP80
| `--LeptopodidaeSP80
| |--LeptosaldinaeSP80
| | |--Leptosalda chiapensis Cobben 1971P92
| | `--Saldolepta Schuh & Polhemus 1980SP80
| | `--S. kistnerorumSP80
| `--LeptopodinaeSP80
| |--Patapius spinosusSP80
| |--Valleriola wilsoniSP80, CGW91
| |--Erianotus lanosusSP80
| |--MartiniolaSP80
| |--Leotichius [Leotichiidae]SP80
| | `--L. speluncarumSP80
| `--LeptopusRD77
| |--L. echinopsL49
| |--L. lanosusL49
| |--L. littoralisG20
| `--L. marmoratusRD77
`--SaldoideaSP80
|--Aepophilus [Aepophilidae]SP80
| `--A. bonnaireiSP80
`--Saldidae [Acanthiidae]SP80
| i. s.: Micracanthia ripulaSP80
| Ioscytus nastiD62
| TeoleucaD62
| LampracanthiaD62
| Oreokora Drake 1962 [=Oreocora]D62
| |--*O. chilensis (Blanchard 1852) (see below for synonymy)D62
| |--O. amazona (Drake 1955) [=Pentacora amazona]D62
| |--O. andensis (Distant 1893) [=Acanthia andensis; incl. Pentacora bruesi Drake 1949]D62
| |--O. angusta (Drake & Carvalho 1948) [=Pentacora angusta]D62
| |--O. bacayana (Drake 1955) [=Pentacora bacayana]D62
| |--O. bergi (Haglund 1899) [=Salda bergi; incl. Saldula lynchi Drake & Carvalho 1948]D62
| |--O. doeringi (Drake & Carvalho 1948) [=Saldula doeringi]D62
| |--O. perula (Drake 1955) [=Pentacora perula]D62
| |--O. pillaona (Drake 1955)D62
| |--O. regilla (Drake 1955) [=Pentacora regilla]D62
| |--O. rogeri (Kirkaldy 1899) [=Acanthia rogeri]D62
| `--O. sola (Drake & Carvalho 1948) [=Saldula sola]D62
| Pelachoris Drake 1962D62
| `--*P. leucographa [=Pentacora leucographa]D62
|--Mesolygaeus [Enicocoridae, Enicocorinae, Mesolygaeidae, Xishanidae]ZG05
| `--M. laiyangensis Ping 1928ZG05
|--ChiloxanthinaeSP80
| |--EnalosaldaSP80
| `--+--ParalosaldaSP80
| `--+--ChiloxanthusSP80
| `--PentacoraSP80
| |--P. salinaD62
| `--P. sphacelata (Uhler 1877) (see below for synonymy)D62
`--SaldinaeSP80
| i. s.: SaldulaCGW91
| |--S. brevicornisCGW91
| |--S. hirsutaD62
| |--S. orthochilaRD77
| |--S. pallipesSP80
| |--S. palustris (Douglas 1874) [=Salda palustris; incl. Saldula fernaldi Drake 1949]D62
| `--S. pexa Drake 1950 [=S. hirsuta pexa]D62
|--SaldunculaSP80
`--+--SaldoidiniSP80
| |--OrthophrysSP80
| `--SaldoidaD62
`--SaldaD62 [SaldiniSP80]
|--S. coxalis [incl. S. argentina]D62
|--S. exiguaP92
|--S. littoralisRD77
|--S. nobilis [incl. S. nobilis var. reuteri]H01
|--S. pallipesH01
|--S. ripariaL49
`--S. saltatoria (Linnaeus 1758)L49, L58 (see below for synonymy)
*Oreokora chilensis (Blanchard 1852) [=Acanthia chilensis, Salda chilensis; incl. A. araucanica Kirkaldy 1889]D62
Pentacora sphacelata (Uhler 1877) [=Salda sphacelata, Saldula sphacelata; incl. Salda rubromaculata Heidemann 1901, Pentacora rubromaculata]D62
Salda saltatoria (Linnaeus 1758)L49, L58 [=Cimex (Oblongus) saltatoriusL49, L58; incl. S. cinctaL49, Acanthia maculataL49, S. zosteraeL49]
*Type species of generic name indicated
References
[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).
[D62] Drake, C. J. 1962. Synonymic data and two new genera of shore-bugs (Hemiptera: Saldidae). Proceedings of the Biological Society of Washington 75: 115–124.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.
[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis revised 10th ed. vol. 1. Laurentii Salvii: Holmiae.
[L49] Lucas, H. 1849. Exploration Scientifique de l’Algérie pendant les années 1840, 1841, 1842 publiée par ordre du gouvernement et avec le concours d’une commission académique. Sciences physiques. Zoologie. II. Histoire naturelle des animaux articulés. Troisième partie. Insectes (suite). Imprimerie Nationale: Paris.
[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.
[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.
[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.
[SP80] Schuh, R. T., & J. T. Polhemus. 1980. Analysis of taxonomic congruence among morphological, ecological and biogeographic data sets for the Leptopodomorpha (Hemiptera). Systematic Zoology 29 (1): 1–26.
[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815+Heteroptera Latreille, 1810). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 143–157. Kluwer Academic Publishers: Dordrecht.
[WEE70] Woodward, T. E., J. W. Evans & V. F. Eastop. 1970. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 387–457. Melbourne University Press.
[ZG05] Zhang, J., V. B. Golub, Y. A. Popov & D. E. Shcherbakov. 2005. Ignotingidae fam. nov. (Insecta: Heteroptera: Tingoidea), the earliest lace bugs from the Upper Mesozoic of eastern China. Cretaceous Research 26: 783–792.