Myrmica

 European fire ant Myrmica rubra, photographed by Alex Wild.

Belongs within: Myrmicinae.

Myrmica is a genus of generalist ants found in Eurasia and North America. The mesosoma retains a moderately distinct mesoepinotal suture and is armed with spines or teeth on the epinotum (Wheeler & Wheeler 1990). Notable species include the European fire ant Myrmica rubra, which is native to Europe but has also become widely introduced in northern North America.

Characters (Wheeler & Wheeler 1990): Antenna with twelve segments. Lateral portion of clypeus not raised; antennal fossa opening onto clypeus without elevated boundary; psammophore absent. Epinotum armed with spines or teeth; promesonotal suture absent on mesosomal dorsum; mesoepinotal suture moderately impressed. Middle and hind tibial spurs finely pectinate. Postpetiole attached to anterior end of first gastric somite, gaster about equally convex above and below.

<==Myrmica Latreille 1804 (see below for synonymy)Z01
    |--*M. rubra (Linnaeus 1758)Z01 [=Formica rubraZ01, Atta (Myrmica) rubraG20]
    |--M. algirica Lucas 1848E12
    |--M. breviceps Smith 1878 (n. d.)B03
    |--M. caeca Jerdon 1851 (n. d.)B03
    |--M. colaxWW90
    |--M. consternens Walker 1859 (n. d.)B03
    |--M. cursor Smith 1878 (n. d.)B03
    |--M. gallieniiAV04
    |--M. humilis Smith 1858 (n. d.)B03
    |--M. inezae Forel 1902B03
    |--M. kurokiiAV04
    |    |--M. k. kurokiiAV04
    |    `--M. k. tipunaAV04
    |--M. laevinodisMS01
    |--M. lobicornisAV04
    |--M. luctuosa Smith 1878 (n. d.)B03
    |--M. margaritae Emery 1889B03
    |--M. mexicanaWW90
    |--M. microrubra Seifert 1993S09
    |--M. obscurata Motschulsky 1863 (n. d.)B03
    |--M. pallinodis Motschulsky 1863 (n. d.)B03
    |--M. ritae Emery 1889B03
    |--M. rugifrons Smith 1858 (n. d.)B03
    |--M. ruginodisAV04
    |--M. rugiventrisWW90
    |--M. rugosa Mayr 1865B03
    |    |--M. r. rugosaAV04
    |    `--M. r. arisanaAV04
    |--M. sabuletiES12
    |--M. scabrinodisES12
    |--M. schenckiAV04
    |--M. slovacaES12
    |--M. smythiesi Forel 1902 (see below for synonymy)B03
    |--M. specioidesES12
    |--M. spinosiorES12
    |--M. striolagasterBS06
    |--M. sulcinodisAV04
    |--M. tahoensisBS06
    |--M. testaceopilosa Lucas 1848E12
    `--M. vandeliES12

Myrmica Latreille 1804 [incl. Dodecamyrmica Arnold 1968, Paramyrmica Cole 1957, Sifolinia Emery 1907, Sommimyrma Menozzi 1925, Symbiomyrma Arnoldi 130]Z01

Myrmica smythiesi Forel 1902 [incl. M. smythiesi var. carbonaria non M. (Monomorium) carbonaria Smith 1858, M. smythiesi var. rupestris]B03

*Type species of generic name indicated

References

[AV04] Als, T. D., R. Vila, N. P. Kandul, D. R. Nash, S.-H. Yen, Y.-F. Hsu, A. A. Mignault, J. J. Boomsma & N. E. Pierce. 2004. The evolution of alternative parasitic life histories in large blue butterflies. Nature 432: 386–390.

[B03] Bingham, C. T. 1903. The Fauna of British India, including Ceylon and Burma. Hymenoptera vol. 2. Ants and Cuckoo-Wasps. Taylor and Francis: London.

[BS06] Brady, S. G., T. R. Schultz, B. L. Fisher & P. S. Ward. 2006. Evaluating alternative hypotheses for the early evolution and diversification of ants. Proceedings of the National Academy of Sciences of the USA 103 (48): 18172–18177.

[ES12] Espadaler, X., & S. Santamaria. 2012. Ecto- and endoparasitic fungi on ants from the Holarctic region. Psyche 2012: 168478.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[S09] Seifert, B. 2009. Cryptic species in ants (Hymenoptera: Formicidae) revisited: we need a change in the alpha-taxonomic approach. Myrmecological News 12: 149–166.

[WW90] Wheeler, G. C., & J. Wheeler. 1990. Insecta: Hymenoptera Formicidae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1277–1294. John Wiley & Sones: New York.

[Z01] Zhou, S. 2001. Ants of Guangxi. Guangxi Normal University Press: Guilin (China).

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