Belongs within: Gamasina.
The Ologamasidae are a family of predatory mites found worldwide, though most diverse in the Southern Hemisphere. Some species are found in intertidal habitats such as members of the genera Hydrogamasus and Periseius. The species Hydrogamasus antarcticus and H. racovitzai are found in tussocks and mosses on the Antarctic Peninsula (Lindquist et al. 2009). Gamasellus racovitzai is a known predator of intertidal oribatids. Species of Hydrogamasus have holodorsal shields in both sexes with the ventro-anal shield fused with the holodorsal shield posteriorly (Evans & Till 1979).
Phoresy is rare among ologamasids but some species of the genera Epiphis and Iphidosoma are associated with carabid beetles. Members of the genera Euryparasitus and Cyrtolaelaps are commonly found in the nests of small mammals (Lindquist et al. 2009). Cyrtolaelaps is characterised by having some of the setae on the dorsal shields distinctly pilose or serrated. Species of Euryparasitus have the tibia and tarsus of the second legs fused in the males (Evans & Till 1979).
Characters (Luxton 1986): Dorsum of adult with holonotal shield or two separate shields; peritrematal shield usually fused with exopodal IV shield, often also with ventro-anal shield.
<==Ologamasidae [Epiphidinae, Euryparasitidae, Gamasellidae, Gamasellini, Gamasiphidae, Ologamasinae]
|--Gamaselliphis Ryke 1961FH93
|--Oriflamella lutulenta Halliday 2008C10
|--Gamasitus Womersley 1956H98
| `--G. obscurus Womersley 1956H98
|--Neogamasellevans Loots & Ryke 1967H98
| `--N. berlesei (Womersley 1956) [=Queenslandolaelaps berlesei]H98
|--Pyriphis Lee 1970H98
| `--P. pyrenoides (Lee 1966) [=Ologamasus pyrenoides]H98
|--Queenslandolaelaps Womersley 1956H98
| `--Q. vitzthumi Womersley 1956H98
|--Solugamasus Lee 1973H98
| `--S. mustela Lee 1973H98
|--Litogamasus Lee 1970L86, L90
| |--*L. setosus (Kramer 1898) [=Cyrtolaelaps setosus]L90
| `--L. falcipes Lee & Hunter 1974L90
|--Evanssellus Ryke 1961H98
| |--*E. foliatus Ryke 1961R61
| `--E. medusa (Lee 1967) [=Heterogamasus medusa]H98
|--Stylochirus Canestrini & Canestrini 1882L91 (see below for synonymy)
| |--‘Gamasiphis (*Periphis)’ hemisphaericusB16
| `--S. rarior (Berlese 1916)FH93 [=Gamasiphis (*Epiphis) rariorFH93, B16]
|--Rhodacaroides Willmann 1959L86
| |--R. aegyptiacus Willmann 1959L86
| `--R. minyaspis Lee 1973H98
|--Euryparasitus Oudemans 1902LKW09, FH93 [incl. Eurylaelaps Oudemans 1902FH93]
| |--E. emarginatus (Koch 1839)FH93 (see below for synonymy)
| `--E. calcarator (Banks 1910) [=Gamasus calcarator]FH93
|--Iphidosoma Berlese 1892L91
| |--*I. fimetarium (Müller 1859)L91 [=Gamasoides fimetariumS22]
| `--I. razumovaeLKW09
|--Hiniphis Lee 1970H98
| |--H. bipala Lee 1973H98
| `--H. hinnus Lee 1970H98
|--Laelaptiella Womersley 1956H98
| |--L. anomala Womersley 1956H98
| `--L. mackerrasae (Domrow 1957) [=Gamasiphis mackerrasae]H98
|--Parasitiphis Womersley 1956H98
| |--*P. littoralis Womersley 1956L90
| |--P. aurora Lee 1970L90
| `--P. jeanneli (André 1947) [incl. Hydrogamasus (Austrohydrogamasus) watsoni Hirschmann 1966]SL71
|--Gamasiphoides Womersley 1956H98
| |--G. aitkeni Lee 1970H98
| |--G. macquariensis (Hirschmann 1966) [=Gamasellus (Hydrogamasellus) macquariensis]SL71
| `--G. propinquus (Womersley 1956) [=Gamasiphis propinqua, Gamasiphoides propinqua]H98
|--Euepicrius Womersley 1942H98
| |--E. filamentosus Womersley 1942H98
| |--E. lootsi Lee 1970H98
| `--E. queenslandicus Womersley 1956H98
|--Geogamasus Lee 1970L86
| |--G. fibularis Karg 1976L86
| |--G. howardi Lee 1970H98
| `--G. minimus Lee 1973H98
|--Hydrogamasellus Hirschmann 1966L86
| |--H. antarcticus (Trägårdh 1907) [=Gamasellus (Hydrogamasellus) antarcticus]H66
| |--H. antennatus Karg 1976L86
| `--H. ubatubaensis (Hirschmann 1966) [=Gamasellus (Hydrogamasellus) ubatubaensis]H66
|--Periseius Womersley 1961 [incl. Psammonsella Haq 1965]FH93
| |--P. brasiliensis Hirschmann 1966L86
| |--P. littoraleH83
| `--P. nobskae (Haq 1965)FH93 [=Psammonsella nobskaeFH93; incl. Pe. (Psammonsella) schusteri Hirschmann 1966H66, FH93]
|--Onchogamasus Womersley 1956H98
| |--O. communis Womersley 1956H98
| |--O. pumilio Lee 1970H98
| |--O. quasicurtipilus Lee 1970H98
| `--O. virguncula Lee 1973H98
|--Acugamasus Lee 1970H98
| |--A. cursor Lee 1970H98
| |--A. elachyaspis Lee 1973H98
| |--A. punctatus (Womersley 1942) [=Digamasellus punctatus, Cyrtolaelaps punctatus]H98
| |--A. semipunctatus (Womersley 1942) [=Digamasellus semipunctatus, Cyrtolaelaps semipunctatus]H98
| `--A. watsoni (Hirschmann 1956) [=Gamasellus watsoni]SL71
|--Antennolaelaps Womersley 1956 [incl. Stylogamasus Womersley 1956]H98
| |--A. affinis Womersley 1956H98
| |--A. aremenae Lee 1973H98
| |--A. celox Lee 1973H98
| |--A. convexa (Womersley 1956) [=Stylogamasus convexus]H98
| `--A. testudo Lee 1970H98
|--Athiasella Lee 1973H98
| |--A. australica (Womersley 1942)H98 (see below for synonymy)
| |--A. dentata (Womersley 1942)H98 (see below for synonymy)
| |--A. markmitchelli (Lee 1970) [=Heydeniella markmitchelli]H98
| |--A. relata (Womersley 1942)H98 (see below for synonymy)
| `--A. relicta (Womersley 1942) (see below for synonymy)H98
|--Caliphis Lee 1970H98
| |--C. calvus Lee 1970H98
| |--C. hickmani (Womersley 1956) [=Gamasiphis hickmani]H98
| |--C. novaezelandiae (Womersley 1956) [=Gamasellus (Neogamasiphis) novaezelandiae]SL71
| |--C. queenslandicus (Womersley 1956) [=Gamasiphis queenslandicus]H98
| |--C. schusteri (Hirschmann 1956) [=Gamasellus (Hydrogamasellus) schusteri]SL71
| `--C. tamborinensis (Womersley 1956) [=Gamasiphis hickmani var. tamborinensis]H98
|--Hydrogamasus Berlese 1892FH93
| |--*H. littoralis (Canestrini & Canestrini 1881) [=Gamasus littoralis]ET79
| |--H. antarcticus Womersley 1937LKW09, SL71
| |--H. giardi (Berlese & Trouessart 1889)H66
| |--H. kensleri Luxton 1967L90
| |--H. racovitzaiLKW09
| |--H. salinus (Laboulbene 1851)H66 [=Gamasus salinusE57]
| `--H. vitzthumi Hirschmann 1966H66
|--Cyrtolaelaps Berlese 1887 [incl. Protolaelaps Trägårdh 1912]FH93
| |--*C. mucronatus (Canestrini & Canestrini 1881)ET79 (see below for synonymy)
| |--C. aster (Berl. 1918)S61, S22 [=Gamasellus (Protolaelaps) asterS61]
| |--C. cervusS22
| |--C. cornutusT05
| |--C. hammeniH83
| |--C. minorK91
| `--C. rectus (Berlese 1920) [=Gamasellus (Protolaelaps) rectus; incl. C. (Gamasellus) leggetti Ryke 1962]FH93
|--CymiphisSL71
| |--C. cymosus (Lee 1966)SL71
| |--C. dumosus (Lee 1966)SL71
| |--C. leptosceles (Lee 1966)SL71
| |--C. mansoni (Lee 1966)SL71
| |--C. nucilis (Lee 1966)SL71
| |--C. validus (Lee 1966)SL71
| `--C. watsoni (Hirschmann 1966) [=Gamasiphis watsoni]SL71
|--Gamasiphis Berlese 1904H98 [incl. Heteroiphis Trägårdh 1952FH93, Micriphis Berlese 1914FH93, Neogamasiphis Trägårdh 1952H98]
| |--G. australicus Womersley 1956H98
| |--G. elegantellusT31
| |--G. elongatulusT31
| |--G. femoralis (Banks 1916) [=Cyrtolaelaps femoralis]H98
| |--G. fornicatus Lee 1970H98
| |--G. illotus Fox 1949FH93
| |--G. lenifornicatus Lee 1973H98
| |--G. pilosellusT31
| |--G. pulchellus (Berlese 1887) [=Gamasus pulchellus; incl. Laelaps bermudaensis Ewing 1920]FH93
| |--G. saccus Lee 1973H98
| |--G. setosus Womersley 1956H98
| `--G. uncifer Trägårdh 1931T31
`--Gamasellus Berlese 1892LKW09, H98 [incl. Laelogamasus Berlese 1905FH93]
|--*G. falciger (Canestrini 1881) [=Gamasus falciger]LI89
|--G. alpinus Schweizer 1949S61
|--G. bellavistae Emberson 1967FH93
|--G. concinus (Womersley 1942) [=Digamasellus concina, Cyrtolaelaps concinnus, D. concinnus, Gamasellus concinnus]H98
|--G. cooperi (Womersley 1961) [=Cyrtolaelaps cooperi]H98
|--G. cophinus Lee 1973H98
|--G. discutatus (Lee 1966) [=Ologamasus discutatus]H98
|--G. gressittiH73
|--G. grossi Lee 1973H98
|--G. humosus Ishikawa 1969LI89
|--G. lanceolatus Liang & Ishikawa 1989LI89
|--G. lativentralis Ishikawa 1983LI89
|--G. litoprothrix (Lee 1966) [=Ologamasus litoprothrix]H98
|--G. montanusW88
|--G. nivalis Schweizer 1949S61
|--G. occidentalisKA90
|--G. racovitzai (Trouessart 1903)MSU91, L86 (see below for synonymy)
| |--G. r. racovitzaiMSU91
| `--G. r. neoorcadensisMSU91
|--G. simpliciseta Liang & Ishikawa 1989LI89
|--G. southcotti (Lee 1966) [=Ologamasus southcotti]H98
|--G. spiricornisS61
|--G. tasmanicus (Womersley 1956) [=Digamasellus tasmanicus, Cyrtolaelaps tasmanicus]H98
|--G. tianmuensis Liang & Ishikawa 1989LI89
|--G. tindalei (Lee 1966) [=Ologamasus tindalei]H98
|--G. tragardhi (Womersley 1942) [=Digamasellus tragardhi, Cyrtolaelaps tragardhi]H98
|--G. venustus Ishikawa 1983LI89
|--G. vibrissatus Emberson 1967K98
|--G. virgosus (Lee 1966) [=Ologamasus virgosus]H98
`--G. xini Liang & Ishikawa 1989LI89
Nomen nudum: Gamasellus arcticus Danks 1981FH93
Athiasella australica (Womersley 1942)H98 [=Hydrogamasus australicusH98, Gamasellus (Hydrogamasellus) australicusH98, H66, Heydeniella australicaH98]
Athiasella dentata (Womersley 1942)H98 [=Hydrogamasus dentatusH98, Gamasellus (Hydrogamasellus) dentatusH98, H66, Heydeniella dentataH98]
Athiasella relata (Womersley 1942)H98 [=Hydrogamasus relatusH98, Gamasellus (Hydrogamasellus) relatusH98, H66, Heydeniella relataH98]
Athiasella relicta (Womersley 1942) [=Hydrogamasus relictus, Gamasellus relictus, Heydeniella relicta; incl. Hydrogamasus relictus var. major Womersley 1942]H98
*Cyrtolaelaps mucronatus (Canestrini & Canestrini 1881)ET79 [=Gamasus micronatusET79, Gamasellus (Protolaelaps) mucronatusS61; incl. Asca affinis Oudms. 1902E57, Gamasellus brevispinosus Trägårdh 1910S22, Protolaelaps brevispinosusS22]
Euryparasitus emarginatus (Koch 1839)FH93 [=Gamasus emarginatusFH93; incl. G. setiger Koch 1839FH93, G. terribilis Michael 1886FH93, *Euryparasitus terribilisET79]
Gamasellus racovitzai (Trouessart 1903)MSU91, L86 [=Gamasus racovitzaiL86, Cyrtolaelaps (Gamasellus) racovitzaL86, Hydrogamasellus racovitzaiL86, Parasitus racovitzaiT05; incl. Zercon tuberculatus Tragardh 1907L86]
Stylochirus Canestrini & Canestrini 1882L91 [incl. Epiphis Berlese 1916L91, Megaliphis Willmann 1938FH93, Periphis Berlese 1914FH93, Physallolaelaps Berlese 1908FH93]
*Type species of generic name indicated
References
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