Belongs within: Teloschistales.
The Physciaceae are a family of lichenised fungi associated with green algae, often found growing on nitrogen-rich substrates. A pseudoparenchymatous upper cortex is present in species of the genus Physcia.
Characters (Cannon & Kirk 2007): Thallus crustose, foliose or fruticose. Ascomata apothecial, sessile, occasionally immersed, usually with a distinct margin. Interascal tissue composed of ± branched and anastomosing paraphyses, often with swollen and/or pigmented apices. Asci with a J+ apical cap, with a conical or funnel-shaped more weakly staining apical cushion, usually with a J+ gelatinized outer layer. Ascospores usually septate, usually dark brown with thickened walls and septa. Anamorphs pycnidial.
<==Physciaceae [Pyxineaceae]EB03
|--+--Pyxine Fr. 1825SS09, KC01 [incl. Phragmopyxine Clem. 1909KC01]
| | `--P. subcinereaSS09
| `--Buellia De Not. 1846 (nom. cons.)SS09, KC01 (see below for synonymy)
| |--*B. disciformisE99
| |--B. aethaleaE99
| |--B. frigidaSS09
| |--B. parasema (nom. illeg.)E99
| `--B. pulverulentaKC01
`--+--Anaptychia Körb. 1848SS09, KC01 (see below for synonymy)
| |--A. ciliarisRS99
| |--‘Hagenia’ crinalisH04
| |--A. palmulataSS09
| `--A. runcinataE98
`--+--Physcia (Schreb.) Michx. 1803SS09, KC01 (see below for synonymy)
| |--P. adscendensM08
| |--P. aipoliaE99
| |--P. albata [=Parmelia albata]G94
| |--P. caesia [incl. Parmelia alboplumbea, Pa. propinqua]G94
| |--P. lacinulataM95
| |--P. semipinnataE98
| |--P. tenellaM08
| `--P. tribaciaF33
`--Heterodermia Trevis. 1868SS09, KC01 [=Pseudophyscia Müll. Arg. 1894KC01; incl. Chaudhuria Zahlbr. 1932KC01]
|--H. chondroidea Web. & Awasthi 1971S03
|--H. circinalisJ87
|--H. comosaJ87
|--H. diademataBB05
|--H. japonicaBB05
|--H. leucomelosJ87
| |--H. l. ssp. leucomelosJ87
| `--H. l. ssp. boryiJ87
|--H. lutescensJ87
|--H. obscurataE98
`--H. vulgarisSS09
Physciaceae incertae sedis:
Amandinea Choisy ex Scheid. & Mayrhofer 1993KC01
Australiaena Matzer, Mayrhofer & Elix 1997KC01
Awasthia Essl. 1978KC01
Dermatiscum Nyl. 1867KC01
Dermiscellum Hafellner, Mayrhofer & Poelt 1979KC01
Dimelaena Norman 1853 [incl. Beltraminia Trevis. 1857]KC01
`--D. orienaG94
Diploicia Massal. 1852 [=Buelliomyces Thomas ex Cif. & Tomas. 1953; incl. Diplotomma Massal. 1852 non Flot. 1849]KC01
Diplotomma Flot. 1849 [=Abacina Norman 1853]KC01
Dirinaria (Tuck.) Clem. 1909KC01
|--D. applanata [incl. Parmelia redacta]G94
`--D. picta [incl. Parmelia plumosa]G94
Hafellia Kalb, Mayrhofer & Scheid. 1986KC01
Hyperphyscia Müll. Arg. 1894 [incl. Physciopsis Choisy 1950, Tetramelaena (Trevis.) Dodge 1971]KC01
PhaeorrhizaEB03
Physconia Poelt 1965KC01
|--P. distortaKC01
|--P. perisidiosaE98
`--P. venustaE98
Rinodina (Ach.) Gray 1821 (see below for synonymy)KC01
Rinodinella Mayrhofer & Poelt 1978KC01
Santessonia Hale & Vobis 1978KC01
`--S. namibensisE99
Tornabea Østh. 1980 [=Tornabenia Trevis. 1853 non Massal. 1853, Tornabeniopsis Follmann 1980 (nom. inv.)]KC01
Phaeophyscia Moberg 1977 [incl. Physciella Essl. 1986]KC01
|--P. chloantha [=Physciella chloantha (Ach.) Essl. 1986; incl. Physcia luganensis]M95
|--P. ciliataM95
|--P. confusa Moberg 1984M95
|--P. constipataM95
|--P. decolor (Kashiwadani) Essl. 1978 [=Physcia decolor Kashiwadani 1975]S03
|--P. denigrata (Hue) Moberg 1994 (see below for synonymy)M95
|--P. endococcinaM95
| |--P. e. var. endococcinaM95
| `--P. e. var. endococcinodes (Poelt) Moberg 1995 (see below for synonymy)M95
|--P. hirtuosa (Krempelh.) Esslinger 1978 [=Physcia hirtuosa Krempelh. 1873]M95
|--P. hispidula (Ach.) Esslinger 1978 (see below for synonymy)M95
| |--P. h. var. hispidulaM95
| `--P. h. var. exornatula (Poelt) Moberg 1995 (see below for synonymy)M95
|--P. kairamoiM95
|--P. melanchra (Hue) Hale 1983 (see below for synonymy)P95
|--P. orbicularisM08
|--P. primaria (Poelt) Trass 1981 [=Physcia hispidula ssp. primaria Poelt 1974]M95
|--P. pyrrhophora (Poelt) Awasthi & Joshi 1978 (see below for synonymy)M95
|--P. rubropulchra (Degelius) Esslinger 1978 (see below for synonymy)M95
|--P. sciastraM95
`--P. squarrosa Kashiwadani 1984M95
Mobergia Mayrhofer & Sheard 1992 non Redlich 1899 (ICZN)KC01
Redonia Dodge 1973 non Roualt 1851 (ICZN)KC01
Anaptychia Körb. 1848SS09, KC01 [=Anaptychiomyces Thomas 1939 (nom. inv.)KC01, Lichenoides Hoffm. 1789KC01; incl. Hagenia Eschw. 1824 non Gmel. 1791KC01, Imbricaria (Schreb.) Michx. 1803 non Juss. 1789KC01, Tornabenia Massal. 1853KC01]
Buellia De Not. 1846 (nom. cons.)SS09, KC01 [incl. Anapyrenium Müll. Arg. 1880KC01, Aplotomma Massal. ex Beltr. 1858KC01, Buelliopsis Schneid. 1897KC01, Gassicurtia Fée 1824 (nom. rej.)KC01, Kemmleria Körb. 1861KC01, Lepropinacia Vent. 1799KC01, Mannia Trevis. 1857 non Opiz 1829KC01, Mattickiolichen Tomas. & Cif. 1952KC01, Mattickiomyces Cif. & Tomas. 1953KC01, Melanaspicilia Vain. 1909KC01, Monerolechia Trevis. 1857KC01, Samboa Tomas. & Cif. 1952KC01, Samboamyces Cif. & Tomas. 1953KC01, Tetramelas Norman 1853KC01]
Phaeophyscia denigrata (Hue) Moberg 1994 [=Physcia denigrata Hue 1900, Physciella denigrata (Hue) Esslinger 1986; incl. Physcia nepalensis Poelt 1974, Physciella nepalensis (Poelt) Esslinger 1986]M95
Phaeophyscia endococcina var. endococcinodes (Poelt) Moberg 1995 [=Physcia endococcinodes Poelt 1974, Pha. endococcinodes (Poelt) Esslinger 1978; incl. Physcia endococcinodes var. megalospora Poelt 1974, Physcia endococcinodes var. stellata Poelt 1984]M95
Phaeophyscia hispidula (Ach.) Esslinger 1978 [incl. Physcia imbricata Vainio 1921, Pha. imbricata (Vainio) Esslinger 1978]M95
Phaeophyscia hispidula var. exornatula (Poelt) Moberg 1995 [=Physcia setosa var. exornatula Zahlbr. 1930, Pha. exornatula (Zahlbr.) Kashiw. 1984, Physcia setosa ssp. exornatula (Zahlbr.) Poelt 1974; incl. Pha. hispidula ssp. limbata Poelt 1974, Pha. limbata (Poelt) Kashiw. 1984]M95
Phaeophyscia melanchra (Hue) Hale 1983 [=Physcia melanchra Hue 1900, Physciella melanchra (Hue) Esslinger 1986; incl. Physcia melanchra f. cupulifera Hue 1900]P95
Phaeophyscia pyrrhophora (Poelt) Awasthi & Joshi 1978 [=Physcia pyrrhophora Poelt 1974; incl. Physcia endococcina var. latiloba Magn. in Awasthi 1957]M95
Phaeophyscia rubropulchra (Degelius) Esslinger 1978 [=Physcia orbicularis f. rubropulchra Degelius 1940, Physcia rubropulchra (Degelius) Moberg 1974; incl. Physcia saxatilis Kashiwadani 1975]M95
Physcia (Schreb.) Michx. 1803SS09, KC01 [incl. Neophyscia Choisy 1959 (n. n.)KC01, Physciomyces Thomas ex Cif. & Tomas. 1953KC01, Psora Hoffm. 1789 (nom. rej.) non Hoffm. 1796 (nom. cons.)KC01]
Rinodina (Ach.) Gray 1821 [incl. Berengeria Trevis. 1853, Courtoisia Marchand 1830, Dictyorinis Clem. 1909, Malmia Choisy 1931, Merorinis Clem. 1909, Mischoblastia Massal. 1852, Mischolecia Choisy 1931, Pachysporaria (Malme) Choisy 1949, Placothallia Trevis. 1853, Pleorinis Clem. 1909, Pseudobuellia de Lesd. 1907, Rhinodinomyces Thomas ex Cif. & Tomas. 1953]KC01
*Type species of generic name indicated
References
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[E99] Eriksson, O. E. (ed.) 1999. Notes on ascomycete systematics. Nos 2440–2755. Myconet 2: 1–41.
[EB03] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, G. Rambold & T. Laessøe (eds.) 2003. Outline of Ascomycota—2003. Myconet 9: 1–89.
[E98] Etayo, J. 1998. Some hypocrealean lichenicolous fungi from southwest Europe. Nova Hedwigia 67 (3–4): 499–509.
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