Belongs within: Helicoidei.
The Polygyrinae are North American snails in which the penis is a rather simple tube without a sheath (Burch & Pearce 1990).
<==Polygyrinae [Polygyrini, Protogona]
|--Inflectarius Pilsbry 1939BP90
| |--I. edentatus (Sampson 1889)NC01
| |--I. inflectus [=Mesodon (Inflectarius) inflectus]BP90
| |--I. magazinensis [=Mesodon (Inflectarius) magazinensis]BP90
| `--I. smithi [=Mesodon (Inflectarius) smithi]BP90
|--Polygyra Say 1818 [Polygyrina, Polygyrinai]BR05
| | i. s.: P. albolabrisCM07
| | P. auriformisP65
| | P. cereolusR64
| | |--P. c. cereolusW65
| | `--P. c. carpenterianusW65
| | P. dorfuellianaP65
| | P. leporinaH68
| | P. planaJ65
| | P. plicataH68
| | P. pustuloidesH68
| | P. texasianaBP90
| | P. thyroidesKC01
| | P. uvuliferaR64
| | |--P. u. uvuliferaR64
| | `--P. u. striataR64
| |--P. (Polygyra)BR17
| | |--*P. (P.) septemvolva (Say 1818) [=Helix septemvolva]BR17
| | | |--P. s. septemvolvaHSD65
| | | |--P. s. febigeriHSD65
| | | `--P. s. volvoxisDH65
| | |--P. (P.) acutedentata [=Helix (P.) acutedentata; incl. H. loisa]C64
| | `--P. (P.) ventrosula [=Helix (P.) ventrosula]C64
| `--P. (Daedalochila Beck 1837) [incl. Lobosculum]BP90
| `--P. (D.) pustulaBP90
`--Praticolella Martens 1892BP90
| i. s.: ‘Helix’ albocincta Binney 1837 [=H. albolineata Gld. 1847, H. albozonata Binney 1847]V15
| P. bakeri Vanatta 1915V15
| P. berlandieriana (Moric. 1833) [=Helix (Helocogena) berlandieriana]V15
| P. campiBP90
| P. griseola (Pfeiffer 1841) [=Helix griseola]V15
| P. jejuna (Say 1821) (see below for synonymy)V15
| |--P. j. jejunaV15
| `--P. j. clavis Vanatta 1915V15
| P. lawae (Lewis 1874) [=Helix (Mesodon) lawii, Polygyra lawae]V15
| |--P. l. lawaeV15
| `--P. l. tallulahensis (Pils. 1898) [=Polygyra lawae tallulahensis]V15
| P. pachyloma (Mkc. 1847) [=Helix pachyloma]V15
| ‘Helix’ virginalis Jan. ex Pfr. 1848V15
|--P. (Praticolella) prisca Auffenberg & Portell 1990AP90
|--P. (Farragutia Vanatta 1915)V15
| `--P. (*F.) mobiliana (Lea 1841) [=Helix mobiliana]V15
| |--P. m. mobilianaV15
| `--P. m. floridana Vanatta 1915V15
`--P. (Filapex Pilsbry 1940)AP90
Praticolella jejuna (Say 1821) [=Helix jejuna; incl. Bradybaena pisum Beck 1837 (n. n.), H. splendidula Anton 1839 (n. n.) non Pfeiffer 1846]V15
*Type species of generic name indicated
References
[AP90] Auffenberg, K., & R. W. Portell. 1990. A new fossil land snail (Gastropoda: Pulmonata: Polygyridae) from the Middle Miocene of northern Florida. Nautilus 103 (4): 143–148.
[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.
[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.
[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.
[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.
[CM07] Cokendolpher, J. C., & P. G. Mitov. 2007. Natural enemies. In: Pinto-da-Rocha, R., G. Machado & G. Giribet (eds) Harvestmen: The Biology of Opiliones pp. 339–373. Harvard University Press: Cambridge (Massachusetts).
[DH65] Dundee, D. S., & P. W. Hermann. 1965. Malacological problems: 1. Nautilus 78 (3): 81–83.
[HSD65] Hermann, P. W., B. C. Strickland & D. S. Dundee. 1965. Baton Rouge greenhouse gastropods. Nautilus 78 (4): 131–133.
[H68] Hubricht, L. 1968. The land snails of Mammoth Cave National Park, Kentucky. Nautilus 82 (1): 24–28.
[J65] Jacobson, M. K. 1965. On some land shells of Eleuthera, Bahamas. Nautilus 78 (4): 120–125.
[KC01] Kirk, P. M., P. F. Cannon, J. C. David & J. A. Stalpers. 2001. Ainsworth & Bisby’s Dictionary of the Fungi 9th ed. CAB International: Wallingford (UK).
[NC01] Nekola, J. C., & B. F. Coles. 2001. Systematics and ecology of Gastrocopta (Gastrocopta) rogersensis (Gastropoda: Pupillidae), a new species of land snail from the Midwest of the United States of America. Nautilus 115 (3): 105–114.
[P65] Pratt, W. L., Jr. 1965. Notes on land snail distribution in Texas. Nautilus 78 (4): 142–143.
[R64] Ross, L. T. 1964. The land mollusks of Siesta Key, Sarasota County, Florida. Nautilus 78 (2): 50–52.
[V15] Vanatta, E. G. 1915. Praticolella. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (2): 194–198.
[W65] Webb, G. R. 1965. Matings between Polygyra cereolus carpenterianus and P. septemvolvis. Nautilus 79 (1): 34–35.