Belongs within: Vetigastropoda.
The Ptychomphalidae are a family of gastropods with a deep labral slit known from the Upper Cambrian to the Lower Jurassic (Knight et al. 1960).
Characters (Knight et al. 1960, as Ptychomphalides): Labral slit deep, selenizone flat or concave, commonly bordered by extended cords or flanges.
<==Ptychomphalidae [Ptychomphalides, Ptychomphalinae, Ptychomphalini]BR17
|--Taeniospira Ulrich & Bridge 1931KC60
| `--*T. eminencensis Ulrich & Bridge 1931KC60
|--Oehlertia Perner 1907 [incl. Pleurotobembex Solle 1956]KC60
| `--*O. senilis (Perner 1907) [=Pleurotomaria (*Oehlertia) senilis]KC60
|--Shwedagonia Batten 1956KC60
| `--*S. elegans Batten 1956KC60
|--Tropidostropha Longstaff 1912W63
| `--*T. griffithii (M’Coy 1844) [=Pleurotomaria griffithii]KC60
|--Euzone Koken 1896W63 [=Polyelasma Cossmann 1897KC60]
| `--*E. alauna Koken 1896 [=*Polyelasma alauna]KC60
|--Mourlonopsis Fletcher 1958W63
| `--*M. strzeleckiana (Morris 1845) [=Pleurotomaria strzeleckiana, Mourlonia strzeleckiana]W63
|--Spiroscala Knight 1945KC60
| `--*S. pagoda Knight 1945KC60
|--Luciellina Kittl 1900KC60
| `--*L. contracta Kittl 1900KC60
|--Ptychomphalina Bayle 1885BR17, BR05 [Ptychomphalinacea, Ptychomphalininae, Ptychomphalinini]
| `--*P. striata (Sowerby 1836) [=Pleurotomaria striata]BR17
|--Euconospira Ulrich in Ulrich & Scofield 1897 [incl. Pernotrochus Chronic 1952, Trechmannia Longstaff 1912]KC60
| |--*E. turbiniformis (Meek & Worthen 1861) [=Pleurotomaria turbiniformis]KC60
| `--E. permianaG31
|--Ptychomphalus Agassiz 1837BR05 [=Cochlicarina Brown 1843KC60; incl. Cryptaenia Eudes-Deslongchamps 1864KC60]
| |--*P. compressus (Sowerby 1813) [=Helicina compressa, *Cochlicarina compressa]KC60
| `--P. expansus (Sowerby 1821)TTE93
|--Walnicholsia Fletcher 1958W63
| |--*W. subcancellata (Morris 1845) [=Pleurotomaria subcancellata, P. cancellata Strzelecki 1845 (n. n.)]W63
| |--W. minuta Waterhouse 1963W63
| |--W. pygmaea Fletcher 1958F71
| `--‘Pleurotomaria’ subheterospira Wanner 1942W63
|--Mourlonia de Koninck 1883BR05 (see below for synonymy)
| |--*M. ‘carinata’ (Sowerby 1812) [=Helix carinatus non H. carinata Müller 1774]KC60
| |--M. (Woolnoughia) angulata Dickins 1963W63
| |--M. duni Etheridge 1898F71
| |--M. helicinaeformis Koninck 1877W63
| |--M. humilis (De Koninck 1877) [=Pleurotomaria humilis]E89
| |--M. lyndonensis Dickins 1957W63
| `--M. propinquaG31
`--Platyteichum Campbell 1953W63
|--*P. costatum Campbell 1953W63
|--P. brenensis (Reed 1932) [=Pleurotomaria brenensis]W63
|--P. coniforme (Etheridge 1892) [=Mourlonia coniformis]W63
|--P. johnstonei Dickins 1961W63
|--P. loratum Waterhouse 1963W63
|--P. spirolaxum Waterhouse 1963W63
`--P. waterhousei (Etheridge 1898) [=Mourlonia waterhousei]W63
Nomen nudum: Shwedagonia orientalis Yü in Pan 1985P85
Mourlonia de Koninck 1883BR05 [incl. Cryptaulus Foerste 1923 non Bavay 1903KC60, Eocryptaulina Foerste 1936KC60, Foersteria Tomlin 1929 non Szépligeti 1896KC60, Promourlonia Longstaff 1924KC60, Spiroraphella Grabau 1936KC60; Mourlonides, Mourloniini]
*Type species of generic name indicated
References
[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.
[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.
[E89] Etheridge, R., Jr. 1889. Remarks on fossils of Permo-Carboniferous age, from north-western Australia, in the Macleay Museum. Proceedings of the Linnean Society of New South Wales, series 2, 4 (2): 199–214, pl. 17.
[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.
[G31] Grabau, A. W. 1931. The Permian of Mongolia: A report on the Permian fauna of the Jisu Honguer limestone of Mongolia and its relations to the Permian of other parts of the world. American Museum of Natural History: New York.
[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.
[P85] Pan Y.-T. 1985. Fossil Gastropoda from the Permian of Sichuan, Guizhou, Yunnan and Guangxi. Acta Palaeontologica Sinica 24 (1): 29–37.
[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.
[W63] Waterhouse, J. B. 1963. Permian gastropods of New Zealand. Part 2 – Pleurotomariacea (in part). New Zealand Journal of Geology and Geophysics 6 (2): 115–154.