Belongs within: Opiliones.
Contains: Conomma, Sorensenius.
The Pyramidopidae are a family of short-legged harvestmen found primarily in tropical Africa, though recent phylogenetic analyses have also placed the Central American Jarmilana pecki here.
Characters (Sharma et al. 2011): Small to medium-sized Laniatores; body length 1.5–5.5 mm. Prosoma narrower than opisthosoma, separated by a transverse groove. Ocularium large, conical or domed, at or very close to frontal margin, entirely granular with conical tubercles or single erect horn on top. Opisthosomal portion of scutum (mesotergum) divided by transverse grooves into five areas, each with one or more rows of granules and often larger spines. Sternum narrowed and genital operculum enclosed by oversized fourth coxae; opisthosoma with four free sternites, two fused sternites in a pre-anal plate and anal operculum. Spiracles hidden under posterior margin of coxa IV. Chelicerae not enlarged; basal segment with inflated dorsodistal bulla. Pedipalps armed with megaspines. Trochanter with single ventral spine; femur with three large, and smaller interspersed, ventral spines, one to two mesodistal and a denticulate dorsal side; patella with two mesal spines; tibia with four spines; tarsus with two to three spines on each side; pedipalpal claw stout. Legs relatively short, with leg II/scutum length ratio from 2 to 5; leg IV longer and stouter than leg II. Distitarsus I with two articles and II with three. Tarsal formula commonly 4–5: 5–14: 5: 6. Tarsi III and IV with two simple claws; pseudonychium and scopula absent. Male genitalia with ventral plate undivided and highly variable (from linguiform to almost annular); dorsum and ventral plate of truncus typically with large paired setae; glans shape highly variable, lacking prominent parastylar conductors.
<==PyramidopidaeKP-GP19
|--'Stygnomma’ pecki Goodnight & Goodnight 1977KP-GP19, K03
`--+--ConommaKP-GP19
`--SorenseniusKP-GP19
Pyramidopidae incertae sedis:
Maiorerus Rambla 1993SPG11
`--*M. randoi Rambla 1993SPG11
Neoconomma Özdikmen 2006SPG11 [=Metaconomma Kauri 1985 non Pickard-Cambridge 1905O06]
`--*N. dentipes (Kauri 1985)SPG11 [=*Metaconomma dentipesO06]
Skufia Roewer 1949SPG11
`--*S. parva Roewer 1949SPG11
Opconomma Roewer 1927SPG11
`--*O. hirsuta Roewer 1927SPG11
Aburiplus Roewer 1949SPG11
`--*A. trochanteralis Roewer 1949SPG11
Kwangonia Kauri 1985SPG11
`--*K. feshiensis Kauri 1985SPG11
Micronimba Roewer 1953SPG11
|--*M. bicurvata Roewer 1953SPG11
|--M. concolor Roewer 1953S92
|--M. femoralis Roewer 1958S92
`--M. pulchella Roewer 1958S92
Opconommula Roewer 1949SPG11
`--*O. spinosa (Roewer 1927)SPG11 [=Opconomma spinosaS92]
Proconomma Roewer 1961SPG11
|--*P. kahuzi Roewer 1961SPG11
`--P. crassipalpis Kauri 1985S92
Tonkouinatus Roewer 1953SPG11
`--*T. magnituber Roewer 1953SPG11
Jarmilana pecki (Goodnight & Goodnight 1977)P-GC17
*Type species of generic name indicated
References
[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.
[KP-GP19] Kury, A. B., A. Pérez-González & D. N. Proud. 2019. A new Indo-Malayan family of Grassatores (Arachnida: Opiliones: Laniatores). Invertebrate Systematics 33: 892–906.
[O06] Özdikmen, H. 2006. Nomenclatural changes for some Laniatores (Opiliones) genera: New substitute names and new combinations. Mun. Ent. Zool. 1 (1): 63–68.
[P-GC17] Pérez-González, A., F. S. Ceccarelli, B. G. O. Monte, D. N. Proud, M. B. DaSilva & M. E. Bichuette. 2017. Light from dark: a relictual troglobite reveals a broader ancestral distribution for kimulid harvestmen (Opiliones: Laniatores: Kimulidae) in South America. PLoS One 12 (11): e0187919.
[SPG11] Sharma, P. P., C. E. Prieto & G. Giribet. 2011. A new family of Laniatores (Arachnida: Opiliones) from the Afrotropics. Invertebrate Systematics 25: 143–154.
[S92] Staręga, W. 1992. An annotated check-list of Afrotropical harvestmen, excluding the Phalangiidae (Opiliones). Annals of the Natal Museum 33 (2): 271–336.