Belongs within: Hesperiidae.
The Pyrginae, flats, are a cosmopolitan group of relatively large, robust and fast-flying skippers whose larvae feed on a wide range of dicotyledonous plants (Braby 2000a).
Characters (from Braby 2000a): Antenna more than half length of costa of fore wing; third segment of labial palp erect or porrect, short to medium in length, stout; fore wing with vein M2 straight near origin, arising midway between M1 and M3 or closer to M1 than M3; hind wing with cell normal, termen rounded; male may have costal gold or linear scent-pouch, containing numerous sex-scales, on costa near base of fore wing.
<==Pyrginae [Pyrgidae]J03
|--CapilaJ03
|--TelemiadesJ03
|--Celaenorrhinus Hübner 1919B2000a
| `--C. fritzgaertneri (Balley 1880)FS54
|--Exometoeca Meyrick 1888B2000a
| `--E. nycteris Meyrick 1888B2000a
|--EagrisJ03
| |--E. decolor Mabille 1890M90
| `--E. melancholica Mabille 1890M90
|--AchalarusS72
| |--A. lyciadesS72
| `--A. toxeus (Plötz 1882)FS54
|--EpargyreusS72
| |--E. clarusS72
| `--E. exadeus (Cramer 1782)S55
|--ThorybesS72
| |--T. bathyllusS72
| `--T. pyladesS72
|--Euschemon Doubleday 1846 [Euschemoninae]B2000a
| `--E. rafflesia (Macleay 1826)B2000a
| |--E. r. rafflesiaB2000a
| `--E. r. alba Mabille 1903B2000a
|--ErynnisS72
| |--E. brizoS72
| |--E. funeralisS72
| `--E. tristisS72
|--HeliopetesS72
| |--H. alana (Reakirt 1868)S55
| |--H. arsalte (Linnaeus 1758)S55
| `--H. ericetorumS72
|--Netrocoryne Felder & Felder 1867B2000a
| |--N. repanda Felder & Felder 1867B2000a
| | |--N. r. repandaB2000a
| | `--N. r. expansa Waterhouse 1932B2000a
| `--N. thaddaeus (Hewitson 1876)B2000a
|--AutochtonS72
| |--A. aunus (Fabricius 1781)FS54
| |--A. cellusS72
| |--A. cincta Herrich-Schäffer & Plötz 1882FS54
| `--A. subalbatus (Plötz 1882)FS54
|--PholisoraS72
| |--P. azteca Scudder 1871FS54
| |--P. catullusS72
| |--P. libyaS72
| `--P. orsines (Godman & Salvin 1896)FS54
|--SyrichthusS72
| |--S. alveolusS72
| |--S. alveus [incl. S. alveus var. onopordi]P01
| |--S. malvae [incl. Papilio alceae, P. malvarum]L49
| |--S. onopordiL49
| `--S. therapneL49
|--Tagiades Hübner 1819B2000a
| |--T. flesusB01a
| |--T. japetus (Stoll 1781)B2000a
| | |--T. j. japetusB2000a
| | `--T. j. janetta Butler 1870B2000a
| `--T. nestus (Felder 1860)B2000b
| |--T. n. nestusB2000b
| `--T. n. korela Mabille 1891B2000b
|--Chaetocneme Felder 1860B2000a
| |--C. beata (Hewitson 1867)B2000a
| |--C. caristus Hewitson 1867B2000a
| |--C. critomedia (Guérin-Méneville 1831)B2000a
| | |--C. c. critomediaB2000a
| | |--C. ‘sphinterifera’ hydra Evans 1934B2000a
| | `--C. c. sphinterifera (Fruhstorfer 1910)B2000a
| |--C. denitza (Hewitson 1867)B2000a
| `--C. porphyropsis (Meyrick & Lower 1902)B2000a
`--Pyrgus Hübner 1819BR05
|--P. dromusB01a
|--P. elmoS72
|--P. machacosaB01b
|--P. maculatus (Bremer & Grey 1852)I92
|--P. malvae (Linnaeus 1758)I92
|--P. notata (Blanchard 1852)FS54
|--P. orcus (Cramer 1782)S55
|--P. spioB1900
`--P. syrichtus (Fabricius 1775)FS54
|--P. s. syrichtusFS54
`--P. s. montivagus (Reakirt 1866)FS54
*Type species of generic name indicated
References
[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.
[B2000a] Braby, M. F. 2000a. Butterflies of Australia: their identification, biology and distribution vol. 1. CSIRO Publishing: Collingwood (Victoria).
[B2000b] Braby, M. F. 2000b. Butterflies of Australia: their identification, biology and distribution vol. 2. CSIRO Publishing: Collingwood (Victoria).
[B1900] Butler, A. G. 1900. On a second collection of butterflies obtained by Mr. Edward M. de Jersey in Nyasaland. Annals and Magazine of Natural History, series 7, 5: 59–63.
[B01a] Butler, A. G. 1901a. An account of a collection of butterflies made by the Rev. K. St. Aubyn Rogers between Mombasa and the forests of Taveta. Annals and Magazine of Natural History, series 7, 7: 22–35.
[B01b] Butler, A. G. 1901b. An account of a collection of butterflies obtained by Lord Delamere, chiefly at Munisu, near Mount Kenya. Annals and Magazine of Natural History, series 7, 7: 197–203.
[FS54] Franz, E., & H. Schröder. 1954. Tagfalter (Lep. Rhopalocera) aus El Salvador. Senckenbergiana Biologica 35: 75–87.
[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.
[J03] Jong, R. de. 2003. Are there butterflies with Gondwanan ancestry in the Australian region? Invertebrate Systematics 17: 143–156.
[L49] Lucas, H. 1849. Exploration Scientifique de l’Algérie pendant les années 1840, 1841, 1842 publiée par ordre du gouvernement et avec le concours d’une commission académique. Sciences physiques. Zoologie. II. Histoire naturelle des animaux articulés. Troisième partie. Insectes (suite). Imprimerie Nationale: Paris.
[M90] Mabille, P. 1890. Lépidoptères (hespérides) nouveaux (2e partie). Annales de la Société Entomologique de France, 6e série 9: clv–clvi.
[P01] Pável, J. 1901. Lepkék [Lepidopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 171–177. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[S55] Schröder, H. 1955. Eine Falter-Ausbeute aus dem westlichen Bolivien. (Ins. Lepid. Rhopal.) Senckenbergiana Biologica 36: 329–338.
[S72] Shields, O. 1972. Flower visitation records for butterflies (Lepidoptera). Pan-Pacific Entomologist 48 (3): 189–203.