Scirpeae

Belongs within: Cyperaceae.
Contains: Kyllinga, Scirpus, Eriophorum, Carex.

The Scirpeae are a tribe of sedges in which all flowers are hermaphroditic, and the glumes are spirally arranged and all floriferous except the lowest (Healy & Edgar 1980). Members of the tribe are found in wetland habitats, some in standing water. Phylogenetic analysis indicates that the group is paraphyletic to the genus Carex which produces unisexual flowers. Also nested in this clade is Sumatroscirpus, characterised by sheathing fertile prophylls, separated from the glumes by an elongated epipodium, antrorsely barbed perianth bristles, and a tubercle on the fruit (Semmouri et al. 2019).

Desmoschoenus spiralis is a New Zealand species with leaves harshly scabrid on the margins and nerves throughout, and the inflorescence a very contracted panicle (Moore & Edgar 1970). Oreobolopsis is a New World genus with a perianth of scale-like tepals.

<==ScirpeaeSB19
    |  i. s.: KyllingaB78
    |         Desmoschoenus Hook. f. 1853ME70
    |           `--D. spiralis (Rich.) Hook. f. 1853 (see below for synonymy)ME70
    |         LipocarphaB78
    |           |--L. argentea [=Hypaelyptum (l. c. for Hypolytrum) argenteum]B78
    |           |--L. aristulata [=Hemicarpha micrantha var. aristulata]H93
    |           |--L. micrantha [=Hemicarpha micrantha; incl. H. micrantha var. minor]H93
    |           |--L. microcephala [=Hypaelyptum (l. c. for Hypolytrum) microcephalum; incl. Scirpus leptocarpus]B78
    |           `--L. occidentalis [=Hemicarpha occidentalis]H93
    |--CalliscirpusSB19
    |    |--C. brachythrixSB19
    |    `--C. crinigerSB19
    `--+--+--+--Amphiscirpus nevadensisSB19
       |  |  `--+--ZameioscirpusSB19
       |  |     |    |--Z. gaimardioidesSB19
       |  |     |    `--+--Z. atacamensisSB19
       |  |     |       `--Z. muticusSB19
       |  |     `--+--Rhodoscirpus asperSB19
       |  |        `--PhylloscirpusSB19
       |  |             |--P. deserticolaSB19
       |  |             `--+--P. acaulisSB19
       |  |                `--P. boliviensisSB19
       |  `--+--‘Scirpus’ ternatanusSB19
       |     `--+--ScirpusSB19
       |        `--EriophorumSB19
       `--+--+--OreobolopsisSB19
          |  |    |--O. clementisSB19
          |  |    `--+--O. tepaliferaSB19
          |  |       `--+--O. inversaSB19
          |  |          `--‘Trichophorum’ rigidumSB19
          |  |--TrichophorumSB19
          |  |    |--T. cespitosumSB19
          |  |    |--T. pumilumSB19
          |  |    |--+--T. clintoniiSB19
          |  |    |  `--T. planifoliumSB19
          |  |    `--+--T. alpinumSB19
          |  |       `--+--T. dioicumSB19
          |  |          `--T. subcapitatumSB19
          |  `--CypringleaSB19
          |       |--C. analectaSB19
          |       `--C. evadensSB19
          `--+--Sumatroscirpus [Sumatroscirpeae]SB19
             |    |--S. paniculatocorymbosusSB19
             |    `--S. rupestrisSB19
             `--CarexSB19

Desmoschoenus spiralis (Rich.) Hook. f. 1853 [=Isolepis spiralis Rich. 1832, Scirpus spiralis (Rich.) Druce 1917 non Rottb. 1773, S. frondosus Boeck. 1878, Anthophyllum urvillei Steud. 1855]^ME70

*Type species of generic name indicated

References

[B78] Bentham, G. 1878. Flora Australiensis: A description of the plants of the Australian Territory vol. 7. Roxburghiaceae to Filices. L. Reeve & Co.: London.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[ME70] Moore, L. B., & E. Edgar. 1970. Flora of New Zealand vol. 2. Indigenous Tracheophyta: Monocotyledones except Gramineae. A. R. Shearer, Government Printer: Wellington (New Zealand).

[SB19] Semmouri, I., K. Bauters, É. Léveillé-Bourret, J. R. Starr, P. Goetghebeur & I. Larridon. 2019. Phylogeny and systematics of Cyperaceae, the evolution and importance of embryo morphology. Botanical Review 85: 1–39.